Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

Prime Medicine, Inc.

CIK 0001894562PRME· quarter ending 2026-06-30

Shares outstanding

181,376,856

as of 2026-07-31 · 10-Q cover page, filed 2026-08-06

Held by 13F filers

106,659,529

58.8% of shares outstanding

Holders

187

-5 vs 2026-03-31 · 22 new, 27 exited

Value

$393.5M

shares +8.2% vs prior quarter

Share classes (cover page of the 10-Q filed 2026-08-06, as of 2026-07-31)

Class	Listed	Shares outstanding	Held by 13F filers	% of class	Holders	CUSIP
Common stock	PRME · NASDAQ	181,376,856	106,659,529	58.8%	187	74168J101

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine

Count	CUSIP	Class (as filers write it)	Holders	Shares held	Class shares	% of class	Value	Options	Mapped via
	74168J101	COM	187	106,659,529	181,376,856	58.8%	$393.5M	3 call / 3 put	13DG	holders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings.

Top holders (187 of 187 · 74168J101)

change vs 2026-03-31

#	Filer	Shares	% of class	Value	Change
1	Alphabet Inc.	16,562,498	9.13%	$61.1M	—
2	BlackRock, Inc.	9,602,041	5.29%	$35.4M	+1,100,333 (+12.9%)
3	ARK Investment Management LLC	8,348,746	4.60%	$30.8M	+2,101,919 (+33.6%)
4	STATE STREET CORP	7,576,460	4.18%	$28.0M	+1,231,335 (+19.4%)
5	T. Rowe Price Investment Management, Inc.	5,237,332	2.89%	$19.3M	-244,561 (-4.5%)
6	VANGUARD CAPITAL MANAGEMENT LLC	4,522,640	2.49%	$16.7M	+48,478 (+1.1%)
7	Balyasny Asset Management L.P.	3,969,082	2.19%	$14.6M	-154,283 (-3.7%)
8	GOLDMAN SACHS GROUP INC	3,430,518	1.89%	$12.7M	+2,460,093 (+253.5%)
9	Casdin Capital, LLC	3,167,175	1.75%	$11.7M	—
10	Woodline Partners LP	3,003,904	1.66%	$11.1M	+591 (+0.0%)
11	UBS Group AG	2,794,689	1.54%	$10.3M	-93,729 (-3.2%)
12	GEODE CAPITAL MANAGEMENT, LLC	2,712,446	1.50%	$10.0M	-31,254 (-1.1%)
13	Sumitomo Mitsui Trust Group, Inc.	2,412,984	1.33%	$8.9M	-527,809 (-17.9%)
14	Amova Asset Management Americas, Inc.	2,407,018	1.33%	$8.9M	-533,775 (-18.2%)
15	CITADEL ADVISORS LLC	2,054,543	1.13%	$7.6M	+1,718,185 (+510.8%)
16	ADAR1 Capital Management, LLC	1,997,552	1.10%	$7.4M	+24,270 (+1.2%)
17	FMR LLC	1,761,884	0.97%	$6.5M	-7,592 (-0.4%)
18	TWO SIGMA INVESTMENTS, LP	1,524,511	0.84%	$5.6M	-48,300 (-3.1%)
19	Redmile Group, LLC	1,389,632	0.77%	$5.1M	-201,200 (-12.6%)
20	FARALLON CAPITAL MANAGEMENT, L.L.C.	1,240,843	0.68%	$4.6M	—
21	Jump Financial, LLC	1,240,429	0.68%	$4.6M	NEW
22	BANK OF AMERICA CORP /DE/	981,335	0.54%	$3.6M	+404,331 (+70.1%)
23	VANGUARD PORTFOLIO MANAGEMENT LLC	961,826	0.53%	$3.5M	+99,622 (+11.6%)
24	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	927,538	0.51%	$3.4M	+10,326 (+1.1%)
25	NORTHERN TRUST CORP	912,597	0.50%	$3.4M	+25,881 (+2.9%)
26	JPMORGAN CHASE & CO	810,095	0.45%	$3.0M	-34,059 (-4.0%)
27	Greenwich Wealth Management LLC	768,001	0.42%	$2.8M	+18,000 (+2.4%)
28	VANGUARD FIDUCIARY TRUST CO	764,633	0.42%	$2.8M	+47,631 (+6.6%)
29	MORGAN STANLEY	708,963	0.39%	$2.6M	+69,506 (+10.9%)
30	Alessandra Capital Management, LLC	666,603	0.37%	$2.5M	+241,093 (+56.7%)
31	D. E. Shaw & Co., Inc.	648,940	0.36%	$2.4M	-744,681 (-53.4%)
32	SUSQUEHANNA INTERNATIONAL GROUP, LLP	648,479	0.36%	$2.4M	-67,350 (-9.4%)
33	Point72 Asset Management, L.P.	533,553	0.29%	$2.0M	NEW
34	MOORE CAPITAL MANAGEMENT, LP	500,000	0.28%	$1.8M	—
35	CHELSEA COUNSEL CO	449,060	0.25%	$1.7M	-13,280 (-2.9%)
36	MILLENNIUM MANAGEMENT LLC	427,097	0.24%	$1.6M	NEW
37	HighTower Advisors, LLC	340,365	0.19%	$1.3M	+110 (+0.0%)
38	DIMENSIONAL FUND ADVISORS LP	304,678	0.17%	$1.1M	+156 (+0.1%)
39	WELLINGTON MANAGEMENT GROUP LLP	304,444	0.17%	$1.1M	+26,324 (+9.5%)
40	Nuveen, LLC	275,931	0.15%	$1.0M	—
41	WELLS FARGO & COMPANY/MN	251,747	0.14%	$929K	-67,803 (-21.2%)
42	Bank of New York Mellon Corp	246,344	0.14%	$909K	-20,875 (-7.8%)
43	IQ EQ FUND MANAGEMENT (IRELAND) Ltd	225,799	0.12%	$833K	+20,410 (+9.9%)
44	MARSHALL WACE, LLP	216,841	0.12%	$800K	+74,884 (+52.8%)
45	XTX Topco Ltd	215,708	0.12%	$796K	+58,426 (+37.1%)
46	Jain Global LLC	215,218	0.12%	$794K	NEW
47	JUPITER ASSET MANAGEMENT LTD	209,490	0.12%	$773K	NEW
48	Swiss National Bank	208,300	0.11%	$769K	+29,700 (+16.6%)
49	PEAK6 LLC	208,057	0.11%	$768K	NEW
50	CITIGROUP INC	195,664	0.11%	$722K	+148,432 (+314.3%)
51	Green Alpha Advisors, LLC	185,023	0.10%	$683K	-6,148 (-3.2%)
52	Keystone Financial Group	166,510	0.09%	$614K	+500 (+0.3%)
53	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	162,338	0.09%	$600K	+39,875 (+32.6%)
54	PRICE T ROWE ASSOCIATES INC /MD/	162,325	0.09%	$599K	+28,570 (+21.4%)
55	HRT FINANCIAL LP	161,510	0.09%	$595K	-451,259 (-73.6%)
56	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	157,049	0.09%	$580K	+70,576 (+81.6%)
57	Bollard Group LLC	151,136	0.08%	$558K	-210,000 (-58.1%)
58	ALLIANCEBERNSTEIN L.P.	150,300	0.08%	$523K	+27,840 (+22.7%)
59	Rafferty Asset Management, LLC	145,275	0.08%	$536K	-70,307 (-32.6%)
60	Invesco Ltd.	140,308	0.08%	$518K	+43,433 (+44.8%)
61	Concurrent Investment Advisors, LLC	131,722	0.07%	$486K	+57,545 (+77.6%)
62	ExodusPoint Capital Management, LP	118,464	0.07%	$437K	NEW
63	AQR CAPITAL MANAGEMENT LLC	117,998	0.07%	$435K	-436,937 (-78.7%)
64	Virtus Investment Advisers, LLC	117,909	0.07%	$435K	+51,401 (+77.3%)
65	RENAISSANCE TECHNOLOGIES LLC	117,412	0.06%	$433K	-623,627 (-84.2%)
66	RHUMBLINE ADVISERS	116,754	0.06%	$431K	+2,485 (+2.2%)
67	FIL Ltd	115,206	0.06%	$425K	—
68	DEUTSCHE BANK AG\	114,472	0.06%	$422K	-42,968 (-27.3%)
69	Steward Partners Investment Advisory, LLC	108,290	0.06%	$400K	-9,166 (-7.8%)
70	Palumbo Wealth Management LLC	103,801	0.06%	$383K	+1,555 (+1.5%)
71	MILESTONE ASSET MANAGEMENT, LLC	100,035	0.06%	$369K	NEW
72	TD Asset Management Inc	94,148	0.05%	$347K	—
73	LPL Financial LLC	92,174	0.05%	$340K	+25,750 (+38.8%)
74	Rockefeller Capital Management L.P.	90,735	0.05%	$335K	-666 (-0.7%)
75	BARCLAYS PLC	88,058	0.05%	$325K	+24,285 (+38.1%)
76	Squarepoint Ops LLC	88,029	0.05%	$325K	-6,037 (-6.4%)
77	COMMONWEALTH EQUITY SERVICES, LLC	87,310	0.05%	$322K	+70,873 (+431.2%)
78	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	85,300	0.05%	$315K	—
79	LOS ANGELES CAPITAL MANAGEMENT LLC	82,305	0.05%	$304K	+29,690 (+56.4%)
80	PROFUND ADVISORS LLC	72,624	0.04%	$268K	-23,182 (-24.2%)
81	Legal & General Group Plc	66,444	0.04%	$245K	—
82	BNP PARIBAS FINANCIAL MARKETS	66,101	0.04%	$244K	+31,688 (+92.1%)
83	BRIDGEWAY CAPITAL MANAGEMENT, LLC	64,800	0.04%	$239K	-70,000 (-51.9%)
84	Jupiter Topco LLC	60,900	0.03%	$224K	NEW
85	Engineers Gate Manager LP	59,133	0.03%	$218K	-51,215 (-46.4%)
86	VANGUARD ASSET MANAGEMENT, Ltd	59,021	0.03%	$218K	—
87	SG Americas Securities, LLC	58,922	0.03%	$217K	NEW
88	&PARTNERS	57,460	0.03%	$211K	-2,010 (-3.4%)
89	MetLife Investment Management, LLC	56,192	0.03%	$207K	+11,984 (+27.1%)
90	Russell Investments Group, Ltd.	50,994	0.03%	$188K	-13,279 (-20.7%)
91	AMERIPRISE FINANCIAL INC	49,941	0.03%	$184K	+357 (+0.7%)
92	Mutual Advisors, LLC	45,000	0.02%	$166K	-200 (-0.4%)
93	Vanguard Global Advisers, LLC	44,269	0.02%	$163K	—
94	Quadrature Capital Ltd	43,663	0.02%	$161K	-3,230 (-6.9%)
95	IvyRock Asset Management (HK) Ltd	41,800	0.02%	$154K	—
96	JONES FINANCIAL COMPANIES LLLP	40,867	0.02%	$140K	+560 (+1.4%)
97	Corient Private Wealth LP	39,078	0.02%	$144K	-2,143 (-5.2%)
98	State of New Jersey Common Pension Fund D	33,569	0.02%	$124K	+2,724 (+8.8%)
99	Cambridge Investment Research Advisors, Inc.	32,854	0.02%	$122K	+20,517 (+166.3%)
100	Crestwood Advisors Group, LLC	32,041	0.02%	$118K	-333 (-1.0%)
101	Police & Firemen's Retirement System of New Jersey	30,751	0.02%	$113K	—
102	OPPENHEIMER & CO INC	29,650	0.02%	$109K	-4,006 (-11.9%)
103	RAYMOND JAMES FINANCIAL INC	29,184	0.02%	$108K	+17,140 (+142.3%)
104	SEI INVESTMENTS CO	29,183	0.02%	$108K	—
105	China Universal Asset Management Co., Ltd.	28,745	0.02%	$106K	-3,288 (-10.3%)
106	ARIZONA STATE RETIREMENT SYSTEM	28,646	0.02%	$106K	+3,855 (+15.5%)
107	ENVESTNET ASSET MANAGEMENT INC	28,278	0.02%	$104K	-762 (-2.6%)
108	MANUFACTURERS LIFE INSURANCE COMPANY, THE	27,353	0.02%	$101K	-1,519 (-5.3%)
109	Vanguard Investments Australia, Ltd.	26,124	0.01%	$96K	—
110	BLUE BELL PRIVATE WEALTH MANAGEMENT, LLC	25,830	0.01%	$95K	-1,185 (-4.4%)
111	Mariner, LLC	25,389	0.01%	$94K	-53,009 (-67.6%)
112	Abacus Planning Group, Inc.	25,188	0.01%	$93K	—
113	INTECH INVESTMENT MANAGEMENT LLC	24,363	0.01%	$90K	-6,175 (-20.2%)
114	Creative Planning	21,295	0.01%	$79K	-6,951 (-24.6%)
115	Magnetar Financial LLC	21,174	0.01%	$78K	+5,390 (+34.1%)
116	E Fund Management Co., Ltd.	20,484	0.01%	$76K	-6,615 (-24.4%)
117	Schonfeld Strategic Advisors LLC	20,189	0.01%	$74K	NEW
118	VOYA INVESTMENT MANAGEMENT LLC	20,138	0.01%	$74K	+2,793 (+16.1%)
119	Vanguard Personalized Indexing Management, LLC	20,083	0.01%	$74K	-1,871 (-8.5%)
120	SHELTON WEALTH MANAGEMENT, LLC	19,105	0.01%	$70K	-961 (-4.8%)
121	CIBC WORLD MARKET INC.	18,662	0.01%	$69K	NEW
122	Graham Capital Management, L.P.	18,516	0.01%	$68K	+2,501 (+15.6%)
123	KLP KAPITALFORVALTNING AS	18,500	0.01%	$68K	—
124	ProShare Advisors LLC	18,243	0.01%	$67K	-4,725 (-20.6%)
125	Zurcher Kantonalbank (Zurich Cantonalbank)	18,225	0.01%	$67K	—
126	IEQ CAPITAL, LLC	16,503	0.01%	$61K	-57,510 (-77.7%)
127	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	15,096	0.01%	$57K	-22,585 (-59.9%)
128	LEVIN CAPITAL STRATEGIES, L.P.	15,000	0.01%	$55K	—
129	PRUDENTIAL FINANCIAL INC	14,755	0.01%	$54K	—
130	THOMPSON DAVIS & CO., INC.	14,590	0.01%	$54K	-2,650 (-15.4%)
131	Sanctuary Advisors, LLC	13,400	0.01%	$49K	-494 (-3.6%)
132	JTC Employer Solutions Trustee Ltd	13,249	0.01%	$49K	—
133	DRW Securities, LLC	13,106	0.01%	$48K	-27,858 (-68.0%)
134	Prime Capital Investment Advisors, LLC	12,945	0.01%	$48K	—
135	Brevan Howard Capital Management LP	12,910	0.01%	$48K	NEW
136	STATE OF WISCONSIN INVESTMENT BOARD	12,681	0.01%	$47K	NEW
137	Pictet Asset Management Holding SA	12,107	0.01%	$45K	—
138	Sequoia Financial Advisors, LLC	11,686	0.01%	$43K	+13 (+0.1%)
139	TEACHER RETIREMENT SYSTEM OF TEXAS	11,440	0.01%	$42K	+382 (+3.5%)
140	Wealthfront Advisers LLC	11,301	0.01%	$42K	+922 (+8.9%)
141	SYON CAPITAL LLC	11,077	0.01%	$41K	NEW
142	VICTORY CAPITAL MANAGEMENT INC	10,979	0.01%	$41K	+162 (+1.5%)
143	BENJAMIN EDWARDS INC	10,329	0.01%	$38K	-3,523 (-25.4%)
144	Forum Financial Management, LP	10,000	0.01%	$37K	—
145	LANARK FINANCIAL, INC.	10,000	0.01%	$37K	+5,000 (+100.0%)
146	Kingsview Wealth Management, LLC	10,000	0.01%	$37K	-25,000 (-71.4%)
147	Federation des caisses Desjardins du Quebec	9,500	0.01%	$35K	—
148	OSAIC HOLDINGS, INC.	9,398	0.01%	$35K	+5,301 (+129.4%)
149	NEW YORK STATE COMMON RETIREMENT FUND	8,434	0.00%	$31K	—
150	Ameritas Investment Partners, Inc.	8,407	0.00%	$31K	—
151	Mirae Asset Global Investments Co., Ltd.	8,078	0.00%	$30K	-1,337 (-14.2%)
152	Tower Research Capital LLC (TRC)	6,683	0.00%	$25K	-15,131 (-69.4%)
153	ROYAL BANK OF CANADA	5,853	0.00%	$22K	-3,401 (-36.8%)
154	Glen Eagle Advisors, LLC	5,000	0.00%	$18K	-5,000 (-50.0%)
155	SIGNATUREFD, LLC	5,000	0.00%	$18K	—
156	NISA INVESTMENT ADVISORS, LLC	4,544	0.00%	$17K	-4,802 (-51.4%)
157	GF FUND MANAGEMENT CO. LTD.	3,822	0.00%	$14K	-214 (-5.3%)
158	Owl Creek Wealth Partners LLC	2,615	0.00%	$10K	—
159	Pacer Advisors, Inc.	2,426	0.00%	$9K	NEW
160	Caitong International Asset Management Co., Ltd	1,644	0.00%	$6K	+1,261 (+329.2%)
161	EverSource Wealth Advisors, LLC	1,376	0.00%	$5K	-624 (-31.2%)
162	Allworth Financial LP	1,278	0.00%	$5K	-3,127 (-71.0%)
163	NewEdge Advisors, LLC	1,000	0.00%	$4K	—
164	Arax Advisory Partners	961	0.00%	$4K	+731 (+317.8%)
165	GAMMA Investing LLC	586	0.00%	$2K	-61 (-9.4%)
166	SBI Securities Co., Ltd.	580	0.00%	$2K	+189 (+48.3%)
167	Quarry LP	540	0.00%	$2K	NEW
168	Sterling Capital Management LLC	536	0.00%	$2K	-2,926 (-84.5%)
169	Fairman Group, LLC	500	0.00%	$2K	—
170	ROTHSCHILD INVESTMENT LLC	384	0.00%	$1K	+2 (+0.5%)
171	PNC Financial Services Group, Inc.	319	0.00%	$1K	—
172	Fortitude Family Office, LLC	308	0.00%	$1K	-128 (-29.4%)
173	Basecamp Wealth Advisors LLC	292	0.00%	$1K	+5 (+1.7%)
174	Parkside Financial Bank & Trust	285	0.00%	$1K	—
175	Optiver Holding B.V.	260	0.00%	$959	-632 (-70.9%)
176	EASTERN BANK	134	0.00%	$494	NEW
177	Covestor Ltd	105	0.00%	$0	+74 (+238.7%)
178	Sachetta, LLC	100	0.00%	$369	NEW
179	HARBOUR INVESTMENTS, INC.	78	0.00%	$288	+4 (+5.4%)
180	Farther Finance Advisors, LLC	55	0.00%	$203	-99 (-64.3%)
181	NFSG Corp	45	0.00%	$166	—
182	NATIXIS	45	0.00%	$166	NEW
183	NATIONAL BANK OF CANADA /FI/	42	0.00%	$155	+32 (+320.0%)
184	Essential Partners LLC	27	0.00%	$100	-981 (-97.3%)
185	IFP Advisors, Inc	23	0.00%	$85	+21 (+1050.0%)
186	Clearstead Advisors, LLC	9	0.00%	$34	NEW
187	Wealth Enhancement Trust Services, Inc.	3	0.00%	$11	NEW

Exited since 2026-03-31 (27 filers; largest 25 shown)

Elevate Capital Advisors, LLC	−151,069
TUDOR INVESTMENT CORP ET AL	−61,546
JANUS HENDERSON GROUP PLC	−60,900
Aquatic Capital Management LLC	−54,230
VARCOV Co.	−41,964
AXQ CAPITAL, LP	−36,742
CWM, LLC	−33,811
TRANSCEND CAPITAL ADVISORS, LLC	−30,070
Turtle Creek Wealth Advisors, LLC	−25,000
NOMURA HOLDINGS INC	−24,823
GROUP ONE TRADING LLC	−24,766
Scientech Research LLC	−17,931
MARTINGALE ASSET MANAGEMENT L P	−14,848
Sterling Investment Advisors, Ltd.	−13,400
MERCER GLOBAL ADVISORS INC /ADV	−12,305
Polymer Capital Management (US) LLC	−10,317
Financial Connections Group, Inc.	−3,712
MTM Investment Management, LLC	−3,090
Scarborough Advisors, LLC	−1,258
MAI Capital Management	−386
Vanguard National Trust Co	−334
True Wealth Design, LLC	−152
Sunbelt Securities, Inc.	−80
Keating Financial Advisory Services, Inc.	−59
US BANCORP \DE\	−33

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting person	Form	Filed	Event	Shares	% of class
T. Rowe Price Investment Management, Inc.	SCHEDULE 13G/A	2026-02-17	2025-12-31	5,995,183	3.3%
GV 2023 GP, L.P.	SCHEDULE 13G/A	2025-11-12	2025-09-30	3,200,000	2.4%
GV 2023 GP, L.L.C.	SCHEDULE 13G/A	2025-11-12	2025-09-30	3,200,000	2.4%
XXVI Holdings Inc.	SCHEDULE 13G/A	2025-11-12	2025-09-30	16,562,498	12.3%
Alphabet Holdings LLC	SCHEDULE 13G/A	2025-11-12	2025-09-30	16,562,498	12.3%
GV 2023, L.P.	SCHEDULE 13G/A	2025-11-12	2025-09-30	3,200,000	2.4%
GV 2021 GP, L.L.C.	SCHEDULE 13G/A	2025-11-12	2025-09-30	3,262,440	2.4%
GV 2019, L.P.	SCHEDULE 13G/A	2025-11-12	2025-09-30	10,100,058	7.5%
Alphabet Inc.	SCHEDULE 13G/A	2025-11-12	2025-09-30	16,562,498	12.3%
GV 2021, L.P.	SCHEDULE 13G/A	2025-11-12	2025-09-30	3,262,440	2.4%
GV 2021 GP, L.P.	SCHEDULE 13G/A	2025-11-12	2025-09-30	3,262,440	2.4%
GV 2019 GP, L.P.	SCHEDULE 13G/A	2025-11-12	2025-09-30	10,100,058	7.5%
GV 2019 GP, L.L.C.	SCHEDULE 13G/A	2025-11-12	2025-09-30	10,100,058	7.5%
FMR LLC	SCHEDULE 13G/A	2025-11-05	2025-09-30	6,731,796	3.8%
Abigail P. Johnson	SCHEDULE 13G/A	2025-11-05	2025-09-30	6,731,796	3.8%
ARCH Venture Partners XII, L.P.	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
Kristina Burow	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Partners X, L.P.	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Partners X, LLC	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Partners XII, LLC	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Fund X Overage, L.P.	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
Keith Crandell	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Partners X Overage, L.P.	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Fund XII, L.P.	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
ARCH Venture Fund X, L.P.	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
Steven Gillis	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,486,894	10.4%
Robert Nelsen	SCHEDULE 13D/A	2025-08-11	2025-08-07	18,587,036	10.4%
Newpath Partners, L.P.	SCHEDULE 13G/A	2025-02-14	2024-12-31	6,105,679	4.7%
Thomas Cahill	SCHEDULE 13G/A	2025-02-14	2024-12-31	6,105,679	4.7%
Newpath Partners GP, L.P.	SCHEDULE 13G/A	2025-02-14	2024-12-31	6,105,679	4.7%
Newpath Partners GP, LLC	SCHEDULE 13G/A	2025-02-14	2024-12-31	6,105,679	4.7%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.