Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

Aura Biosciences, Inc.

CIK 0001501796AURA· quarter ending 2026-06-30

Shares outstanding

103,704,809

as of 2026-08-06 · 10-Q cover page, filed 2026-08-11

Held by 13F filers

98,336,907

94.8% of shares outstanding

Holders

146

+20 vs 2026-03-31 · 36 new, 16 exited

Value

$698.2M

shares +93.3% vs prior quarter

Share classes (cover page of the 10-Q filed 2026-08-11, as of 2026-08-06)

Class	Listed	Shares outstanding	Held by 13F filers	% of class	Holders	CUSIP
Common stock	AURA · NASDAQ	103,704,809	98,336,907	94.8%	146	05153U107

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine

Count	CUSIP	Class (as filers write it)	Holders	Shares held	Class shares	% of class	Value	Options	Mapped via
	05153U107	COM	146	98,336,907	103,704,809	94.8%	$698.2M	1 call / 2 put	13DG	holders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings. 1 further CUSIPs map to this company (mostly bonds) but no 13F filer held them this quarter.

Top holders (146 of 146 · 05153U107)

change vs 2026-03-31

#	Filer	Shares	% of class	Value	Change
1	TCG Crossover Management, LLC	9,533,333	9.19%	$67.7M	NEW
2	MONTANOVA CAPITAL, LLC	7,206,217	6.95%	$51.2M	+2,833,333 (+64.8%)
3	Frazier Life Sciences Management, L.P.	7,100,000	6.85%	$50.4M	+2,000,000 (+39.2%)
4	Nantahala Capital Management, LLC	5,902,060	5.69%	$41.9M	+3,784,363 (+178.7%)
5	Long Focus Capital Management, LLC	5,119,582	4.94%	$36.3M	+500,000 (+10.8%)
6	BlackRock, Inc.	4,707,674	4.54%	$33.4M	+1,121,034 (+31.3%)
7	ADAGE CAPITAL PARTNERS GP, L.L.C.	4,640,000	4.47%	$32.9M	-241,125 (-4.9%)
8	Caligan Partners LP	4,298,248	4.14%	$30.5M	+4,068,384 (+1769.9%)
9	VANGUARD CAPITAL MANAGEMENT LLC	4,100,342	3.95%	$29.1M	+1,856,514 (+82.7%)
10	FCPM III SERVICES B.V.	4,053,775	3.91%	$28.8M	NEW
11	ORBIMED ADVISORS LLC	3,333,333	3.21%	$23.7M	NEW
12	Medicxi Ventures Management (Jersey) Ltd	3,039,892	2.93%	$21.6M	—
13	Jupiter Topco LLC	2,500,000	2.41%	$17.8M	NEW
14	RA CAPITAL MANAGEMENT, L.P.	2,500,000	2.41%	$17.8M	NEW
15	Balyasny Asset Management L.P.	2,354,619	2.27%	$16.7M	+1,796,965 (+322.2%)
16	FRANKLIN RESOURCES INC	2,072,726	2.00%	$14.7M	—
17	GEODE CAPITAL MANAGEMENT, LLC	1,754,721	1.69%	$12.5M	+443,859 (+33.9%)
18	EcoR1 Capital, LLC	1,666,667	1.61%	$11.8M	NEW
19	Redmile Group, LLC	1,666,667	1.61%	$11.8M	NEW
20	Trails Edge Capital Partners, LP	1,443,333	1.39%	$10.2M	NEW
21	Foresite Capital Management VI LLC	1,333,334	1.29%	$9.5M	NEW
22	STATE STREET CORP	1,222,984	1.18%	$8.7M	+258,314 (+26.8%)
23	Woodline Partners LP	1,210,183	1.17%	$8.6M	NEW
24	Bellevue Group AG	1,166,667	1.12%	$8.3M	NEW
25	Vivo Capital, LLC	1,166,667	1.12%	$8.3M	NEW
26	ACORN CAPITAL ADVISORS, LLC	929,077	0.90%	$6.6M	+333,333 (+56.0%)
27	BOOTHBAY FUND MANAGEMENT, LLC	906,356	0.87%	$6.4M	+885,954 (+4342.5%)
28	VANGUARD PORTFOLIO MANAGEMENT LLC	871,342	0.84%	$6.2M	+480,008 (+122.7%)
29	LEVIN CAPITAL STRATEGIES, L.P.	838,067	0.81%	$6.0M	-103,333 (-11.0%)
30	B Group, Inc.	764,243	0.74%	$5.4M	+166,667 (+27.9%)
31	STEMPOINT CAPITAL LP	714,280	0.69%	$5.1M	NEW
32	VANGUARD FIDUCIARY TRUST CO	637,204	0.61%	$4.5M	+240,044 (+60.4%)
33	DIMENSIONAL FUND ADVISORS LP	618,891	0.60%	$4.4M	+60,733 (+10.9%)
34	MILLENNIUM MANAGEMENT LLC	617,467	0.60%	$4.4M	-480,160 (-43.7%)
35	AWM Investment Company, Inc.	591,285	0.57%	$4.2M	NEW
36	NORTHERN TRUST CORP	549,462	0.53%	$3.9M	+118,940 (+27.6%)
37	Regency Capital Management Inc.\DE	540,395	0.52%	$3.8M	+7,900 (+1.5%)
38	GOLDMAN SACHS GROUP INC	464,668	0.45%	$3.3M	+285,942 (+160.0%)
39	PRICE T ROWE ASSOCIATES INC /MD/	278,471	0.27%	$2.0M	-136,720 (-32.9%)
40	FMR LLC	255,331	0.25%	$1.8M	+252,437 (+8722.8%)
41	MORGAN STANLEY	174,692	0.17%	$1.2M	-49,349 (-22.0%)
42	RENAISSANCE TECHNOLOGIES LLC	174,200	0.17%	$1.2M	+72,100 (+70.6%)
43	Integral Health Asset Management, LLC	170,000	0.16%	$1.2M	NEW
44	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	163,156	0.16%	$1.2M	+66,017 (+68.0%)
45	XTX Topco Ltd	149,931	0.14%	$1.1M	+60,410 (+67.5%)
46	WELLINGTON MANAGEMENT GROUP LLP	132,122	0.13%	$938K	+4,579 (+3.6%)
47	Quantinno Capital Management LP	130,656	0.13%	$928K	+39,581 (+43.5%)
48	ADAR1 Capital Management, LLC	130,152	0.13%	$924K	NEW
49	Bank of New York Mellon Corp	118,593	0.11%	$842K	+19,160 (+19.3%)
50	PLATINUM INVESTMENT MANAGEMENT LTD	111,865	0.11%	$794K	—
51	AQR CAPITAL MANAGEMENT LLC	109,889	0.11%	$780K	+63,746 (+138.1%)
52	Qube Research & Technologies Ltd	109,564	0.11%	$778K	NEW
53	BNP PARIBAS FINANCIAL MARKETS	106,351	0.10%	$755K	+16,296 (+18.1%)
54	BANK OF AMERICA CORP /DE/	87,162	0.08%	$619K	-38,194 (-30.5%)
55	Nuveen, LLC	83,804	0.08%	$595K	+11,092 (+15.3%)
56	Fisher Asset Management, LLC	83,508	0.08%	$593K	—
57	Jump Financial, LLC	73,000	0.07%	$518K	NEW
58	MARTINGALE ASSET MANAGEMENT L P	71,258	0.07%	$506K	NEW
59	UBS Group AG	69,230	0.07%	$492K	-49,674 (-41.8%)
60	D. E. Shaw & Co., Inc.	67,405	0.06%	$479K	NEW
61	RHUMBLINE ADVISERS	63,657	0.06%	$452K	-1,063 (-1.6%)
62	Permanent Capital Management, LP	62,369	0.06%	$443K	—
63	TBH Global Asset Management, LLC	60,322	0.06%	$428K	—
64	WATER ISLAND CAPITAL LLC	60,000	0.06%	$426K	NEW
65	ALLIANCEBERNSTEIN L.P.	52,760	0.05%	$353K	+11,100 (+26.6%)
66	CITADEL ADVISORS LLC	51,902	0.05%	$369K	+39,399 (+315.1%)
67	Virtus Investment Advisers, LLC	51,045	0.05%	$362K	NEW
68	PRUDENTIAL FINANCIAL INC	49,050	0.05%	$348K	NEW
69	Invesco Ltd.	44,850	0.04%	$318K	+10,582 (+30.9%)
70	AMERIPRISE FINANCIAL INC	42,682	0.04%	$303K	+413 (+1.0%)
71	DEUTSCHE BANK AG\	41,844	0.04%	$297K	+3,873 (+10.2%)
72	WELLS FARGO & COMPANY/MN	40,918	0.04%	$291K	+23,924 (+140.8%)
73	BANK OF NOVA SCOTIA	40,768	0.04%	$289K	NEW
74	Vanguard Global Advisers, LLC	37,682	0.04%	$268K	-2,107 (-5.3%)
75	MANUFACTURERS LIFE INSURANCE COMPANY, THE	36,038	0.03%	$256K	-21,381 (-37.2%)
76	Jain Global LLC	30,600	0.03%	$217K	NEW
77	COMMONWEALTH EQUITY SERVICES, LLC	30,316	0.03%	$215K	+32 (+0.1%)
78	Quadrature Capital Ltd	29,872	0.03%	$212K	+19,205 (+180.0%)
79	LETKO, BROSSEAU & ASSOCIATES INC	28,300	0.03%	$201K	—
80	JPMORGAN CHASE & CO	27,382	0.03%	$188K	+3,847 (+16.3%)
81	CANADA PENSION PLAN INVESTMENT BOARD	27,000	0.03%	$192K	—
82	MetLife Investment Management, LLC	25,220	0.02%	$179K	+258 (+1.0%)
83	China Universal Asset Management Co., Ltd.	24,554	0.02%	$174K	+13,285 (+117.9%)
84	CITIGROUP INC	23,432	0.02%	$166K	+10,950 (+87.7%)
85	Engineers Gate Manager LP	22,233	0.02%	$158K	NEW
86	nVerses Capital, LLC	20,798	0.02%	$148K	NEW
87	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	20,663	0.02%	$147K	-206 (-1.0%)
88	TD Asset Management Inc	19,996	0.02%	$142K	-20,506 (-50.6%)
89	Russell Investments Group, Ltd.	17,990	0.02%	$128K	+14,315 (+389.5%)
90	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	17,940	0.02%	$123K	+5,020 (+38.9%)
91	Point72 Asset Management, L.P.	17,864	0.02%	$127K	-3,844 (-17.7%)
92	Inscription Capital, LLC	17,806	0.02%	$126K	—
93	Creative Planning	15,472	0.01%	$110K	+1,399 (+9.9%)
94	BARCLAYS PLC	14,462	0.01%	$103K	+2,922 (+25.3%)
95	BROWN ADVISORY INC	13,400	0.01%	$95K	—
96	ExodusPoint Capital Management, LP	13,278	0.01%	$94K	-4,677 (-26.0%)
97	NEW YORK STATE COMMON RETIREMENT FUND	13,104	0.01%	$93K	—
98	PRELUDE CAPITAL MANAGEMENT, LLC	13,000	0.01%	$92K	—
99	INTECH INVESTMENT MANAGEMENT LLC	11,521	0.01%	$82K	-4,260 (-27.0%)
100	MERCER GLOBAL ADVISORS INC /ADV	11,440	0.01%	$81K	NEW
101	Police & Firemen's Retirement System of New Jersey	11,397	0.01%	$81K	—
102	Vontobel Holding Ltd.	11,000	0.01%	$78K	-4,000 (-26.7%)
103	Polymer Capital Management (US) LLC	10,284	0.01%	$73K	NEW
104	Focus Partners Wealth	10,049	0.01%	$71K	NEW
105	VOYA INVESTMENT MANAGEMENT LLC	10,006	0.01%	$71K	NEW
106	Tower Research Capital LLC (TRC)	6,952	0.01%	$49K	+3,039 (+77.7%)
107	Parallel Advisors, LLC	5,665	0.01%	$40K	—
108	Mirae Asset Global Investments Co., Ltd.	4,624	0.00%	$33K	+1,312 (+39.6%)
109	Ameritas Investment Partners, Inc.	4,544	0.00%	$32K	—
110	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	4,517	0.00%	$32K	—
111	Arax Advisory Partners	4,422	0.00%	$31K	+3,400 (+332.7%)
112	Legal & General Group Plc	4,096	0.00%	$29K	—
113	ROYAL BANK OF CANADA	3,121	0.00%	$23K	+1,741 (+126.2%)
114	NISA INVESTMENT ADVISORS, LLC	2,543	0.00%	$18K	NEW
115	Rockefeller Capital Management L.P.	2,439	0.00%	$17K	-430 (-15.0%)
116	GF FUND MANAGEMENT CO. LTD.	2,043	0.00%	$15K	+627 (+44.3%)
117	Smartleaf Asset Management LLC	1,790	0.00%	$12K	—
118	GAMMA Investing LLC	1,047	0.00%	$7K	+187 (+21.7%)
119	Versant Capital Management, Inc	1,023	0.00%	$7K	-438 (-30.0%)
120	LOS ANGELES CAPITAL MANAGEMENT LLC	820	0.00%	$6K	-12,454 (-93.8%)
121	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	618	0.00%	$4K	-1,759 (-74.0%)
122	Allworth Financial LP	502	0.00%	$4K	-243 (-32.6%)
123	FIFTH THIRD BANCORP	488	0.00%	$3K	—
124	Farther Finance Advisors, LLC	434	0.00%	$3K	+426 (+5325.0%)
125	SIGNATUREFD, LLC	365	0.00%	$3K	-157 (-30.1%)
126	US BANCORP \DE\	292	0.00%	$2K	+212 (+265.0%)
127	Optiver Holding B.V.	288	0.00%	$2K	-104 (-26.5%)
128	OSAIC HOLDINGS, INC.	210	0.00%	$1K	-716 (-77.3%)
129	IFP Advisors, Inc	209	0.00%	$1K	+14 (+7.2%)
130	Sterling Capital Management LLC	205	0.00%	$1K	-916 (-81.7%)
131	Parkside Financial Bank & Trust	169	0.00%	$1K	—
132	Quarry LP	147	0.00%	$1K	NEW
133	EverSource Wealth Advisors, LLC	138	0.00%	$980	-23 (-14.3%)
134	SJS Investment Consulting Inc.	130	0.00%	$923	+128 (+6400.0%)
135	Essential Partners LLC	116	0.00%	$824	-431 (-78.8%)
136	True Wealth Design, LLC	87	0.00%	$618	+73 (+521.4%)
137	Blue Trust, Inc.	86	0.00%	$611	—
138	PNC Financial Services Group, Inc.	83	0.00%	$589	—
139	MassMutual Private Wealth & Trust, FSB	78	0.00%	$554	—
140	Caitong International Asset Management Co., Ltd	71	0.00%	$504	+69 (+3450.0%)
141	ROTHSCHILD INVESTMENT LLC	68	0.00%	$483	+37 (+119.4%)
142	Silver Grove Financial Group, Inc.	41	0.00%	$291	NEW
143	SBI Securities Co., Ltd.	34	0.00%	$241	—
144	Clearstead Advisors, LLC	23	0.00%	$165	NEW
145	Fortitude Family Office, LLC	7	0.00%	$50	NEW
146	Inspire Investing, LLC	5	0.00%	$38	—

Exited since 2026-03-31 (16 filers; largest 16 shown)

Matrix Capital Management Company, LP	−6,922,870
Maven Securities LTD	−50,000
AMERICAN CENTURY COMPANIES INC	−48,323
Brevan Howard Capital Management LP	−29,746
Susquehanna Portfolio Strategies, LLC	−28,729
SUSQUEHANNA INTERNATIONAL GROUP, LLP	−27,483
EntryPoint Capital, LLC	−26,300
49 WEALTH MANAGEMENT, LLC	−21,704
Trexquant Investment LP	−20,998
BRIDGEWAY CAPITAL MANAGEMENT, LLC	−16,276
SIMPLEX TRADING, LLC	−8,430
CWM, LLC	−4,018
MAI Capital Management	−16
Covestor Ltd	−10
Root Financial Partners, LLC	−7
Advisory Services Network, LLC	−6

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting person	Form	Filed	Event	Shares	% of class
Forbion Growth III Management B.V.	SCHEDULE 13G	2026-10-01	2026-09-25	5,248,841	5.1%
ForGrowth III PA B.V.	SCHEDULE 13G	2026-10-01	2026-09-25	5,248,841	5.1%
Forbion Growth Opportunities Fund III Cooperatief U.A.	SCHEDULE 13G	2026-10-01	2026-09-25	5,248,841	5.1%
JOHN HELMERS	SCHEDULE 13G	2026-08-14	2026-06-30	5,119,582	4.9%
FHMLS XI, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	358,810	0.3%
Frazier Life Sciences Public Fund, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	5,807,260	5.6%
Frazier Life Sciences XII, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	805,410	0.8%
Frazier Life Sciences XI, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	358,810	0.3%
FHMLS X, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	128,520	0.1%
FHMLS X, L.L.C.	SCHEDULE 13G/A	2026-08-14	2026-06-30	128,520	0.1%
FHMLS XI, L.L.C.	SCHEDULE 13G/A	2026-08-14	2026-06-30	358,810	0.3%
FHMLS XII, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	805,410	0.8%
FHMLS XII, L.L.C.	SCHEDULE 13G/A	2026-08-14	2026-06-30	805,410	0.8%
Patrick J. Heron	SCHEDULE 13G/A	2026-08-14	2026-06-30	128,520	0.1%
LONG FOCUS CAPITAL MANAGEMENT, LLC	SCHEDULE 13G	2026-08-14	2026-06-30	5,119,582	4.9%
FHMLSP, L.L.C.	SCHEDULE 13G/A	2026-08-14	2026-06-30	5,807,260	5.6%
FHMLSP, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	5,807,260	5.6%
James N. Topper	SCHEDULE 13G/A	2026-08-14	2026-06-30	128,520	0.1%
Frazier Life Sciences X, L.P.	SCHEDULE 13G/A	2026-08-14	2026-06-30	128,520	0.1%
Phillip Gross	SCHEDULE 13G/A	2026-08-12	2026-06-30	9,104,294	8.44%
Robert Atchinson	SCHEDULE 13G/A	2026-08-12	2026-06-30	9,104,294	8.44%
Adage Capital Management, L.P.	SCHEDULE 13G/A	2026-08-12	2026-06-30	9,104,294	8.44%
BlackRock, Inc.	SCHEDULE 13G/A	2026-07-27	2026-06-30	4,582,436	4.4%
Chen Yu	SCHEDULE 13G	2026-05-12	2026-05-05	9,533,333	9.9%
TCG Crossover Fund III, L.P.	SCHEDULE 13G	2026-05-12	2026-05-05	4,766,666	5%
TCG Crossover Fund II, L.P.	SCHEDULE 13G	2026-05-12	2026-05-05	4,766,667	5%
TCG Crossover GP III, LLC	SCHEDULE 13G	2026-05-12	2026-05-05	4,766,666	5%
TCG Crossover GP II, LLC	SCHEDULE 13G	2026-05-12	2026-05-05	4,766,667	5%
David E. Goel	SCHEDULE 13D/A	2026-05-11	2026-05-07	0	0%
Matrix Capital Management Company LP	SCHEDULE 13D/A	2026-05-11	2026-05-07	0	0%
Rajesh Parekh, Ph.D.	SCHEDULE 13G/A	2026-02-17	2026-02-13	2,071,857	3.335%
Advent-Harrington Impact Fund LP	SCHEDULE 13G/A	2026-02-17	2026-02-13	2,023,857	3.258%
Advent Life Sciences Fund I LP	SCHEDULE 13G/A	2026-02-17	2026-02-13	2,023,857	3.258%
ALS III Carry and Co-Invest LP	SCHEDULE 13G/A	2026-02-17	2026-02-13	2,023,857	3.258%
Advent Life Sciences LLP	SCHEDULE 13G/A	2026-02-17	2026-02-13	2,023,857	3.258%
Advent Life Sciences Fund III LP	SCHEDULE 13G/A	2026-02-17	2026-02-13	2,023,857	3.258%
FHMLSP Overage, L.P.	SCHEDULE 13G/A	2025-08-14	2025-06-30	918,510	1.5%
Daniel Estes	SCHEDULE 13G/A	2025-08-14	2025-06-30	0	0%
FHMLSP Overage, L.L.C.	SCHEDULE 13G/A	2025-08-14	2025-06-30	918,510	1.5%
James Brush	SCHEDULE 13G/A	2025-08-14	2025-06-30	0	0%
Frazier Life Sciences Public Overage Fund, L.P.	SCHEDULE 13G/A	2025-08-14	2025-06-30	918,510	1.5%
Albert Cha	SCHEDULE 13G/A	2025-08-14	2025-06-30	0	0%
Aaron Cowen	SCHEDULE 13G/A	2025-08-13	2025-06-30	4,701,775	7.6%
Suvretta Capital Management, LLC	SCHEDULE 13G/A	2025-08-13	2025-06-30	4,701,775	7.6%
Averill Master Fund, Ltd.	SCHEDULE 13G/A	2025-08-13	2025-06-30	4,137,797	6.6%
Eventide Asset Management, LLC	SCHEDULE 13G/A	2025-05-14	2025-03-31	0	0%
Finny Kuruvilla, M.D. Ph. D.	SCHEDULE 13G/A	2025-05-14	2025-03-31	0	0%
Robin C. John	SCHEDULE 13G/A	2025-05-14	2025-03-31	0	0%
LONG FOCUS CAPITAL MASTER, LTD.	SCHEDULE 13G/A	2025-01-03	2024-12-24	1,782,997	3.6%
CONDAGUA, LLC	SCHEDULE 13G/A	2025-01-03	2024-12-24	724,777	1.4%
GLENN HELMERS	SCHEDULE 13G/A	2025-01-03	2024-12-24	724,777	1.4%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.