Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

Protara Therapeutics, Inc.

CIK 0001359931TARA· quarter ending 2026-06-30

Shares outstanding

59,095,488

as of 2026-08-07 · 10-Q cover page, filed 2026-08-11

Held by 13F filers

46,130,930

78.1% of shares outstanding

Holders

119

+5 vs 2026-03-31 · 25 new, 20 exited

Value

$176.2M

shares +1.3% vs prior quarter

Share classes (cover page of the 10-Q filed 2026-08-11, as of 2026-08-07)

Class	Listed	Shares outstanding	Held by 13F filers	% of class	Holders	CUSIP
Common stock	TARA · NASDAQ	59,095,488	46,130,930	78.1%	119	74365U107

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine

Count	CUSIP	Class (as filers write it)	Holders	Shares held	Class shares	% of class	Value	Options	Mapped via
	74365U107	COM STK	119	46,130,930	59,095,488	78.1%	$176.2M	0 call / 1 put	13DG	holders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings.

Top holders (119 of 119 · 74365U107)

change vs 2026-03-31

#	Filer	Shares	% of class	Value	Change
1	Jupiter Topco LLC	4,696,709	7.95%	$17.9M	NEW
2	BlackRock, Inc.	3,663,499	6.20%	$14.0M	+268,531 (+7.9%)
3	MILLENNIUM MANAGEMENT LLC	3,101,166	5.25%	$11.8M	+269,958 (+9.5%)
4	Velan Capital Investment Management LP	2,844,652	4.81%	$10.9M	+10,000 (+0.4%)
5	VANGUARD CAPITAL MANAGEMENT LLC	2,300,133	3.89%	$8.8M	+181,115 (+8.5%)
6	Blackstone Inc.	2,265,514	3.83%	$8.7M	—
7	ORBIMED ADVISORS LLC	2,220,000	3.76%	$8.5M	NEW
8	Sio Capital Management, LLC	1,921,872	3.25%	$7.3M	-47,967 (-2.4%)
9	ACORN CAPITAL ADVISORS, LLC	1,752,115	2.96%	$6.7M	—
10	Integral Health Asset Management, LLC	1,500,000	2.54%	$5.7M	-250,000 (-14.3%)
11	5AM Venture Management, LLC	1,369,782	2.32%	$5.2M	—
12	GEODE CAPITAL MANAGEMENT, LLC	1,326,246	2.24%	$5.1M	+149,046 (+12.7%)
13	CITADEL ADVISORS LLC	1,232,365	2.09%	$4.7M	+163,807 (+15.3%)
14	STATE STREET CORP	1,068,295	1.81%	$4.1M	+63,153 (+6.3%)
15	Qube Research & Technologies Ltd	1,045,516	1.77%	$4.0M	+868,512 (+490.7%)
16	Balyasny Asset Management L.P.	1,026,691	1.74%	$3.9M	NEW
17	BAKER BROS. ADVISORS LP	971,614	1.64%	$3.7M	—
18	ADAR1 Capital Management, LLC	940,512	1.59%	$3.6M	—
19	BANK OF AMERICA CORP /DE/	882,179	1.49%	$3.4M	-43,739 (-4.7%)
20	MARSHALL WACE, LLP	701,021	1.19%	$2.7M	+650,290 (+1281.8%)
21	DIMENSIONAL FUND ADVISORS LP	683,230	1.16%	$2.6M	+151,963 (+28.6%)
22	NORTHERN TRUST CORP	552,306	0.93%	$2.1M	+177,454 (+47.3%)
23	Point72 Asset Management, L.P.	548,932	0.93%	$2.1M	-603,782 (-52.4%)
24	GOLDMAN SACHS GROUP INC	546,227	0.92%	$2.1M	+148,246 (+37.2%)
25	Catalio Capital Management, LP	520,148	0.88%	$2.0M	-59,876 (-10.3%)
26	MORGAN STANLEY	508,434	0.86%	$1.9M	-48,540 (-8.7%)
27	Walleye Capital LLC	505,432	0.86%	$1.9M	+62,031 (+14.0%)
28	VANGUARD PORTFOLIO MANAGEMENT LLC	491,657	0.83%	$1.9M	+79,053 (+19.2%)
29	UBS Group AG	456,773	0.77%	$1.7M	-945,640 (-67.4%)
30	VANGUARD FIDUCIARY TRUST CO	334,683	0.57%	$1.3M	+25,841 (+8.4%)
31	Verdad Advisers, LP	312,249	0.53%	$1.2M	NEW
32	JACOBS LEVY EQUITY MANAGEMENT, INC	295,565	0.50%	$1.1M	+245,310 (+488.1%)
33	RENAISSANCE TECHNOLOGIES LLC	251,500	0.43%	$961K	+125,800 (+100.1%)
34	OPPENHEIMER & CO INC	238,362	0.40%	$911K	+56,797 (+31.3%)
35	AQR CAPITAL MANAGEMENT LLC	208,969	0.35%	$798K	+142,164 (+212.8%)
36	Jump Financial, LLC	203,100	0.34%	$776K	+136,321 (+204.1%)
37	CIBC WORLD MARKET INC.	166,495	0.28%	$636K	-9,000 (-5.1%)
38	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	157,971	0.27%	$603K	+10,083 (+6.8%)
39	Bank of New York Mellon Corp	137,689	0.23%	$526K	+8,018 (+6.2%)
40	J. Goldman & Co LP	135,000	0.23%	$516K	-721,247 (-84.2%)
41	SUPERSTRING CAPITAL MANAGEMENT LP	121,457	0.21%	$464K	—
42	Assenagon Asset Management S.A.	119,908	0.20%	$458K	-447,713 (-78.9%)
43	Fisher Asset Management, LLC	99,566	0.17%	$380K	NEW
44	AMERICAN CENTURY COMPANIES INC	98,487	0.17%	$376K	+5,518 (+5.9%)
45	BRIDGEWAY CAPITAL MANAGEMENT, LLC	85,330	0.14%	$326K	+15,000 (+21.3%)
46	Nuveen, LLC	72,610	0.12%	$277K	+8,750 (+13.7%)
47	TWO SIGMA INVESTMENTS, LP	69,652	0.12%	$266K	+31,814 (+84.1%)
48	FMR LLC	68,200	0.12%	$261K	+11,105 (+19.5%)
49	XTX Topco Ltd	63,948	0.11%	$244K	+5,147 (+8.8%)
50	RHUMBLINE ADVISERS	63,245	0.11%	$242K	+31,170 (+97.2%)
51	TRUIST FINANCIAL CORP	60,000	0.10%	$229K	—
52	Trexquant Investment LP	59,616	0.10%	$228K	+41,683 (+232.4%)
53	DEUTSCHE BANK AG\	57,713	0.10%	$220K	+3,738 (+6.9%)
54	ALPS ADVISORS INC	57,041	0.10%	$218K	+13,012 (+29.6%)
55	AMERIPRISE FINANCIAL INC	56,267	0.10%	$215K	NEW
56	WELLS FARGO & COMPANY/MN	54,596	0.09%	$209K	+23,899 (+77.9%)
57	PRUDENTIAL FINANCIAL INC	50,730	0.09%	$194K	-68,800 (-57.6%)
58	NANO CAP NEW MILLENNIUM GROWTH FUND L P	50,000	0.08%	$191K	+13,500 (+37.0%)
59	SUSQUEHANNA INTERNATIONAL GROUP, LLP	40,380	0.07%	$154K	-76,246 (-65.4%)
60	Alyeska Investment Group, L.P.	39,726	0.07%	$152K	-310,274 (-88.6%)
61	Quadrature Capital Ltd	39,319	0.07%	$150K	NEW
62	PRICE T ROWE ASSOCIATES INC /MD/	38,483	0.07%	$148K	+2,177 (+6.0%)
63	Marex Group Ltd	35,250	0.06%	$135K	-20,000 (-36.2%)
64	WINTON GROUP Ltd	32,800	0.06%	$125K	+4,700 (+16.7%)
65	PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO	29,700	0.05%	$113K	NEW
66	Russell Investments Group, Ltd.	28,469	0.05%	$109K	+25,793 (+963.9%)
67	MetLife Investment Management, LLC	28,378	0.05%	$108K	-398 (-1.4%)
68	Invesco Ltd.	26,856	0.05%	$103K	+8,895 (+49.5%)
69	CITIGROUP INC	24,145	0.04%	$92K	+9,669 (+66.8%)
70	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	24,095	0.04%	$92K	-6 (-0.0%)
71	JPMORGAN CHASE & CO	22,608	0.04%	$94K	+1,118 (+5.2%)
72	Jain Global LLC	21,000	0.04%	$80K	NEW
73	Vanguard Global Advisers, LLC	20,127	0.03%	$77K	-32,472 (-61.7%)
74	PRELUDE CAPITAL MANAGEMENT, LLC	19,613	0.03%	$75K	NEW
75	EntryPoint Capital, LLC	19,286	0.03%	$74K	NEW
76	LPL Financial LLC	19,130	0.03%	$73K	-17,000 (-47.1%)
77	BARCLAYS PLC	18,607	0.03%	$71K	-29,655 (-61.4%)
78	Squarepoint Ops LLC	18,124	0.03%	$69K	NEW
79	ALLIANCEBERNSTEIN L.P.	18,040	0.03%	$94K	NEW
80	IEQ CAPITAL, LLC	17,929	0.03%	$68K	NEW
81	Police & Firemen's Retirement System of New Jersey	17,569	0.03%	$67K	—
82	MANUFACTURERS LIFE INSURANCE COMPANY, THE	14,878	0.03%	$57K	+1,330 (+9.8%)
83	INTECH INVESTMENT MANAGEMENT LLC	13,562	0.02%	$52K	-227 (-1.6%)
84	Mariner, LLC	11,899	0.02%	$45K	NEW
85	VOYA INVESTMENT MANAGEMENT LLC	11,469	0.02%	$44K	NEW
86	D. E. Shaw & Co., Inc.	11,113	0.02%	$42K	NEW
87	COMMONWEALTH EQUITY SERVICES, LLC	11,000	0.02%	$42K	—
88	Corient Private Wealth LP	10,690	0.02%	$41K	NEW
89	Essential Partners LLC	9,802	0.02%	$37K	-70 (-0.7%)
90	NEW YORK STATE COMMON RETIREMENT FUND	7,900	0.01%	$30K	—
91	ROYAL BANK OF CANADA	7,763	0.01%	$29K	+4,263 (+121.8%)
92	BNP PARIBAS FINANCIAL MARKETS	7,536	0.01%	$29K	-3,471 (-31.5%)
93	Tower Research Capital LLC (TRC)	4,990	0.01%	$19K	+387 (+8.4%)
94	Legal & General Group Plc	4,490	0.01%	$17K	—
95	Ameritas Investment Partners, Inc.	3,298	0.01%	$13K	—
96	Arax Advisory Partners	1,671	0.00%	$6K	NEW
97	Sterling Capital Management LLC	1,055	0.00%	$4K	-648 (-38.1%)
98	State of Wyoming	960	0.00%	$4K	NEW
99	GAMMA Investing LLC	732	0.00%	$3K	+155 (+26.9%)
100	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	720	0.00%	$3K	-2,091 (-74.4%)
101	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	451	0.00%	$2K	—
102	JONES FINANCIAL COMPANIES LLLP	402	0.00%	$2K	NEW
103	Allworth Financial LP	371	0.00%	$1K	+288 (+347.0%)
104	SIMPLEX TRADING, LLC	339	0.00%	$1K	-4,110 (-92.4%)
105	EverSource Wealth Advisors, LLC	269	0.00%	$1K	+139 (+106.9%)
106	Aspect Partners, LLC	230	0.00%	$879	+124 (+117.0%)
107	PNC Financial Services Group, Inc.	117	0.00%	$447	-4 (-3.3%)
108	Caitong International Asset Management Co., Ltd	108	0.00%	$413	+106 (+5300.0%)
109	SBI Securities Co., Ltd.	103	0.00%	$393	—
110	OSAIC HOLDINGS, INC.	100	0.00%	$382	—
111	Farther Finance Advisors, LLC	92	0.00%	$351	-25 (-21.4%)
112	EASTERN BANK	71	0.00%	$271	NEW
113	True Wealth Design, LLC	52	0.00%	$199	+28 (+116.7%)
114	ROTHSCHILD INVESTMENT LLC	51	0.00%	$195	+8 (+18.6%)
115	SJS Investment Consulting Inc.	20	0.00%	$77	+12 (+150.0%)
116	GROUP ONE TRADING LLC	14	0.00%	$53	-1,283 (-98.9%)
117	JPL Wealth Management, LLC	4	0.00%	$16	NEW
118	Clearstead Advisors, LLC	3	0.00%	$12	NEW
119	IFP Advisors, Inc	2	0.00%	$8	NEW

Exited since 2026-03-31 (20 filers; largest 20 shown)

JANUS HENDERSON GROUP PLC	−5,207,011
AIGH Capital Management LLC	−1,622,733
Patient Square Capital LP	−1,030,846
Burkehill Global Management, LP	−450,000
ExodusPoint Capital Management, LP	−116,305
FOX RUN MANAGEMENT, L.L.C.	−35,965
Almanack Investment Partners, LLC.	−23,782
Engineers Gate Manager LP	−21,200
BAILARD, INC.	−18,617
AXQ CAPITAL, LP	−13,002
Cerity Partners LLC	−12,748
TORONTO DOMINION BANK	−12,470
Virtu Financial LLC	−10,212
CWM, LLC	−5,846
NISA INVESTMENT ADVISORS, LLC	−4,800
MAI Capital Management	−148
Basecamp Wealth Advisors LLC	−82
Sunbelt Securities, Inc.	−12
Root Financial Partners, LLC	−5
Wealth Preservation Advisors, LLC	−1

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting person	Form	Filed	Event	Shares	% of class
JANUS HENDERSON GROUP Ltd.	SCHEDULE 13G/A	2026-08-13	2026-06-30	4,696,709	8.4%
Millennium Management LLC	SCHEDULE 13G/A	2026-07-07	2026-07-01	3,201,936	5.7%
Millennium Group Management LLC	SCHEDULE 13G/A	2026-07-07	2026-07-01	3,201,936	5.7%
Integrated Core Strategies (US) LLC	SCHEDULE 13G/A	2026-07-07	2026-07-01	2,879,674	5.1%
Israel A. Englander	SCHEDULE 13G/A	2026-07-07	2026-07-01	3,201,936	5.7%
JANUS HENDERSON GROUP PLC	SCHEDULE 13G/A	2026-05-15	2026-03-31	5,207,011	9.6%
BlackRock, Inc.	SCHEDULE 13G	2026-04-27	2026-03-31	3,387,459	6.3%
Janus Henderson Biotech Innovation Master Fund Ltd	SCHEDULE 13G/A	2025-11-14	2025-09-30	3,579,068	9.3%
Rajeev Shah	SCHEDULE 13G/A	2025-08-14	2025-06-30	1,971,765	4.9%
RA Capital Management, L.P.	SCHEDULE 13G/A	2025-08-14	2025-06-30	1,971,765	4.9%
RA Capital Healthcare Fund, L.P.	SCHEDULE 13G/A	2025-08-14	2025-06-30	1,971,765	4.9%
Peter Kolchinsky	SCHEDULE 13G/A	2025-08-14	2025-06-30	1,971,765	4.9%
Phillip Gross	SCHEDULE 13G/A	2025-05-12	2025-03-31	0	0%
Adage Capital Management, L.P.	SCHEDULE 13G/A	2025-05-12	2025-03-31	0	0%
Robert Atchinson	SCHEDULE 13G/A	2025-05-12	2025-03-31	0	0%
Opaleye Management, Inc.	SCHEDULE 13G/A	2025-04-28	2025-03-31	1,730,000	4.71%
Kenneth Griffin	SCHEDULE 13G/A	2025-02-14	2024-12-31	1,627,734	4.7%
Citadel Securities LLC	SCHEDULE 13G/A	2025-02-14	2024-12-31	15,203	0%
Citadel Securities Group LP	SCHEDULE 13G/A	2025-02-14	2024-12-31	15,203	0%
Citadel Securities GP LLC	SCHEDULE 13G/A	2025-02-14	2024-12-31	15,203	0%
Citadel GP LLC	SCHEDULE 13G/A	2025-02-14	2024-12-31	1,612,531	4.6%
Citadel Advisors LLC	SCHEDULE 13G/A	2025-02-14	2024-12-31	1,612,531	4.6%
Citadel Advisors Holdings LP	SCHEDULE 13G/A	2025-02-14	2024-12-31	1,612,531	4.6%
Woodline Partners LP	SCHEDULE 13G/A	2025-02-14	2024-12-31	1,331,258	3.9%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.