Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

CODEXIS, INC.

CIK 0001200375CDXS· quarter ending 2026-06-30

Shares outstanding

109,505,765

as of 2026-08-06 · 10-Q cover page, filed 2026-08-11

Held by 13F filers

69,770,141

63.7% of shares outstanding

Holders

148

+13 vs 2026-03-31 · 26 new, 13 exited

Value

$157.6M

shares +5.0% vs prior quarter

Share classes (cover page of the 10-Q filed 2026-08-11, as of 2026-08-06)

Class	Listed	Shares outstanding	Held by 13F filers	% of class	Holders	CUSIP
Common stock	CDXS · NASDAQ	109,505,765	69,770,141	63.7%	148	192005106

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine

Count	CUSIP	Class (as filers write it)	Holders	Shares held	Class shares	% of class	Value	Options	Mapped via
	192005106	COM	148	69,770,141	109,505,765	63.7%	$157.6M	2 call / 0 put	13DG	holders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings.

Top holders (148 of 148 · 192005106)

change vs 2026-03-31

#	Filer	Shares	% of class	Value	Change
1	BlackRock, Inc.	7,284,352	6.65%	$16.5M	+255,579 (+3.6%)
2	Nantahala Capital Management, LLC	7,247,226	6.62%	$16.4M	+3,123,613 (+75.7%)
3	Opaleye Management Inc.	6,785,000	6.20%	$15.3M	—
4	Aberdeen Group plc	6,769,954	6.18%	$15.3M	-1,910,414 (-22.0%)
5	MILLENNIUM MANAGEMENT LLC	5,549,139	5.07%	$12.5M	+1,551,009 (+38.8%)
6	Telemark Asset Management, LLC	3,846,411	3.51%	$8.7M	—
7	AMERIPRISE FINANCIAL INC	3,809,440	3.48%	$8.6M	+408,944 (+12.0%)
8	VANGUARD CAPITAL MANAGEMENT LLC	3,617,109	3.30%	$8.2M	-51,398 (-1.4%)
9	Assenagon Asset Management S.A.	2,556,412	2.33%	$5.8M	+1,805,242 (+240.3%)
10	GEODE CAPITAL MANAGEMENT, LLC	2,162,393	1.97%	$4.9M	+277,040 (+14.7%)
11	STATE STREET CORP	1,756,814	1.60%	$4.0M	+89,860 (+5.4%)
12	D. E. Shaw & Co., Inc.	1,342,014	1.23%	$3.0M	+362,167 (+37.0%)
13	JACOBS LEVY EQUITY MANAGEMENT, INC	1,330,205	1.21%	$3.0M	+242,178 (+22.3%)
14	VANGUARD PORTFOLIO MANAGEMENT LLC	1,307,171	1.19%	$3.0M	+427,832 (+48.7%)
15	AQR CAPITAL MANAGEMENT LLC	845,431	0.77%	$1.9M	+236,830 (+38.9%)
16	TWO SIGMA INVESTMENTS, LP	696,599	0.64%	$1.6M	+454,644 (+187.9%)
17	NORTHERN TRUST CORP	682,940	0.62%	$1.5M	+25,801 (+3.9%)
18	FRANKLIN RESOURCES INC	678,640	0.62%	$1.5M	-9,570 (-1.4%)
19	MORGAN STANLEY	649,170	0.59%	$1.5M	+176,329 (+37.3%)
20	VANGUARD FIDUCIARY TRUST CO	581,058	0.53%	$1.3M	+30,050 (+5.5%)
21	Man Group plc	565,593	0.52%	$1.3M	NEW
22	Post Resch Tallon Group Inc.	563,478	0.51%	$1.3M	-8,000 (-1.4%)
23	PERKINS CAPITAL MANAGEMENT INC	526,100	0.48%	$1.2M	-1,400 (-0.3%)
24	FMR LLC	478,483	0.44%	$1.1M	+65,886 (+16.0%)
25	GOLDMAN SACHS GROUP INC	434,924	0.40%	$983K	-389,946 (-47.3%)
26	ALGERT GLOBAL LLC	430,910	0.39%	$974K	+233,463 (+118.2%)
27	Nuveen, LLC	392,267	0.36%	$887K	+24,824 (+6.8%)
28	ACADIAN ASSET MANAGEMENT LLC	358,285	0.33%	$804K	+234,536 (+189.5%)
29	TUDOR INVESTMENT CORP ET AL	357,925	0.33%	$809K	-179,908 (-33.5%)
30	DIMENSIONAL FUND ADVISORS LP	313,386	0.29%	$709K	+28,750 (+10.1%)
31	Russell Investments Group, Ltd.	308,095	0.28%	$696K	+249,616 (+426.8%)
32	PLATINUM INVESTMENT MANAGEMENT LTD	264,464	0.24%	$598K	—
33	ROYAL BANK OF CANADA	262,926	0.24%	$594K	+47,561 (+22.1%)
34	Potomac Capital Management, Inc.	260,000	0.24%	$588K	-14,200 (-5.2%)
35	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	242,211	0.22%	$547K	—
36	XTX Topco Ltd	240,627	0.22%	$544K	+73,085 (+43.6%)
37	BANK OF AMERICA CORP /DE/	240,517	0.22%	$544K	+66,562 (+38.3%)
38	UBS Group AG	239,907	0.22%	$542K	-1,943,868 (-89.0%)
39	Bank of New York Mellon Corp	226,039	0.21%	$511K	+16,394 (+7.8%)
40	CITADEL ADVISORS LLC	191,580	0.17%	$433K	-96,015 (-33.4%)
41	Invesco Ltd.	162,271	0.15%	$367K	-18,654 (-10.3%)
42	BARCLAYS PLC	155,274	0.14%	$351K	+181 (+0.1%)
43	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	148,051	0.14%	$335K	NEW
44	ALLIANCEBERNSTEIN L.P.	126,430	0.12%	$206K	+8,300 (+7.0%)
45	BRIDGEWAY CAPITAL MANAGEMENT, LLC	125,908	0.11%	$285K	—
46	STRS OHIO	121,000	0.11%	$273K	+21,900 (+22.1%)
47	JANE STREET GROUP, LLC	120,815	0.11%	$273K	-799,149 (-86.9%)
48	MARSHALL WACE, LLP	117,135	0.11%	$265K	NEW
49	Stonepine Capital Management, LLC	109,662	0.10%	$248K	-308,332 (-73.8%)
50	RHUMBLINE ADVISERS	103,301	0.09%	$233K	+3,987 (+4.0%)
51	WELLS FARGO & COMPANY/MN	101,926	0.09%	$230K	+45,295 (+80.0%)
52	DEUTSCHE BANK AG\	96,229	0.09%	$217K	+3,147 (+3.4%)
53	SUSQUEHANNA INTERNATIONAL GROUP, LLP	82,523	0.08%	$187K	+29,282 (+55.0%)
54	LPL Financial LLC	76,560	0.07%	$173K	+1,500 (+2.0%)
55	PRICE T ROWE ASSOCIATES INC /MD/	66,924	0.06%	$152K	+3,331 (+5.2%)
56	PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO	63,753	0.06%	$144K	—
57	Numerai GP LLC	60,209	0.05%	$136K	NEW
58	PANAGORA ASSET MANAGEMENT INC	58,200	0.05%	$132K	NEW
59	Mariner, LLC	54,637	0.05%	$123K	+31,108 (+132.2%)
60	Cerity Partners LLC	53,322	0.05%	$121K	+20,430 (+62.1%)
61	MetLife Investment Management, LLC	52,011	0.05%	$118K	NEW
62	ENVESTNET ASSET MANAGEMENT INC	51,375	0.05%	$116K	+14,961 (+41.1%)
63	LOS ANGELES CAPITAL MANAGEMENT LLC	48,125	0.04%	$109K	-85,540 (-64.0%)
64	Virtu Financial LLC	46,601	0.04%	$105K	NEW
65	IEQ CAPITAL, LLC	46,265	0.04%	$105K	NEW
66	CITIGROUP INC	43,333	0.04%	$98K	+31,424 (+263.9%)
67	JPMORGAN CHASE & CO	41,861	0.04%	$98K	+454 (+1.1%)
68	GSA CAPITAL PARTNERS LLP	41,811	0.04%	$94K	-75,112 (-64.2%)
69	RENAISSANCE TECHNOLOGIES LLC	41,500	0.04%	$94K	-157,233 (-79.1%)
70	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	39,804	0.04%	$90K	-2,614 (-6.2%)
71	Frec Markets, Inc.	37,269	0.03%	$84K	+1,016 (+2.8%)
72	OSAIC HOLDINGS, INC.	36,366	0.03%	$82K	-199 (-0.5%)
73	COMMONWEALTH EQUITY SERVICES, LLC	36,204	0.03%	$82K	+4,704 (+14.9%)
74	BANK OF MONTREAL /CAN/	34,998	0.03%	$79K	—
75	Krilogy Financial LLC	34,825	0.03%	$79K	-1,432 (-3.9%)
76	Dynamic Technology Lab Private Ltd	33,000	0.03%	$74K	NEW
77	Vanguard Global Advisers, LLC	32,602	0.03%	$74K	—
78	MERCER GLOBAL ADVISORS INC /ADV	32,350	0.03%	$73K	+14,671 (+83.0%)
79	AMERICAN CENTURY COMPANIES INC	30,501	0.03%	$69K	NEW
80	BLAIR WILLIAM & CO/IL	30,219	0.03%	$68K	+201 (+0.7%)
81	EVERGREEN CAPITAL MANAGEMENT LLC	29,000	0.03%	$66K	—
82	Capital Advisors, Ltd. LLC	28,890	0.03%	$65K	—
83	PRUDENTIAL FINANCIAL INC	26,380	0.02%	$60K	+12,800 (+94.3%)
84	GUGGENHEIM CAPITAL LLC	26,158	0.02%	$59K	-792 (-2.9%)
85	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	25,275	0.02%	$61K	+6,416 (+34.0%)
86	MANUFACTURERS LIFE INSURANCE COMPANY, THE	25,205	0.02%	$57K	-2,234 (-8.1%)
87	INTECH INVESTMENT MANAGEMENT LLC	21,454	0.02%	$48K	-2,793 (-11.5%)
88	Police & Firemen's Retirement System of New Jersey	19,441	0.02%	$44K	—
89	VOYA INVESTMENT MANAGEMENT LLC	18,561	0.02%	$42K	+3,745 (+25.3%)
90	Vanguard Personalized Indexing Management, LLC	17,948	0.02%	$41K	-235 (-1.3%)
91	Simplicity Wealth,LLC	17,438	0.02%	$39K	+4,130 (+31.0%)
92	NEW YORK STATE COMMON RETIREMENT FUND	16,888	0.02%	$38K	—
93	Corient Private Wealth LP	16,738	0.02%	$38K	NEW
94	Creative Planning	15,488	0.01%	$35K	NEW
95	BNP PARIBAS FINANCIAL MARKETS	15,461	0.01%	$35K	-5,230 (-25.3%)
96	Navalign, LLC	15,340	0.01%	$35K	—
97	EP Wealth Advisors, LLC	15,277	0.01%	$35K	NEW
98	China Universal Asset Management Co., Ltd.	14,466	0.01%	$33K	-1,563 (-9.8%)
99	ProShare Advisors LLC	14,303	0.01%	$32K	-2,673 (-15.7%)
100	Elmwood Wealth Management, Inc.	12,695	0.01%	$29K	-9,000 (-41.5%)
101	OXFORD ASSET MANAGEMENT LLP	12,095	0.01%	$27K	NEW
102	Graham Capital Management, L.P.	12,074	0.01%	$27K	NEW
103	Tower Research Capital LLC (TRC)	12,055	0.01%	$27K	-1,517 (-11.2%)
104	Abel Hall, LLC	11,916	0.01%	$27K	NEW
105	Caerus Investment Advisors, LLC	11,636	0.01%	$26K	+1,056 (+10.0%)
106	Arax Advisory Partners	11,607	0.01%	$26K	+11,306 (+3756.1%)
107	CAPTRUST FINANCIAL ADVISORS	11,553	0.01%	$26K	NEW
108	Allspring Global Investments Holdings, LLC	11,210	0.01%	$27K	NEW
109	Quadrature Capital Ltd	10,181	0.01%	$23K	-54,018 (-84.1%)
110	GAMCO INVESTORS, INC. ET AL	10,000	0.01%	$23K	—
111	Legal & General Group Plc	9,364	0.01%	$21K	+2,222 (+31.1%)
112	Ameritas Investment Partners, Inc.	8,428	0.01%	$19K	—
113	Pinnacle Wealth Planning Services, Inc.	8,273	0.01%	$19K	—
114	NISA INVESTMENT ADVISORS, LLC	7,497	0.01%	$17K	+7,300 (+3705.6%)
115	FNY Investment Advisers, LLC	5,000	0.00%	$11K	-15,000 (-75.0%)
116	GAMMA Investing LLC	4,766	0.00%	$11K	+360 (+8.2%)
117	PNC Financial Services Group, Inc.	4,646	0.00%	$11K	+770 (+19.9%)
118	GROUP ONE TRADING LLC	4,609	0.00%	$10K	NEW
119	Mirae Asset Global Investments Co., Ltd.	4,065	0.00%	$9K	-646 (-13.7%)
120	Allworth Financial LP	2,304	0.00%	$5K	+794 (+52.6%)
121	Optiver Holding B.V.	2,275	0.00%	$5K	NEW
122	GF FUND MANAGEMENT CO. LTD.	1,913	0.00%	$4K	+105 (+5.8%)
123	EverSource Wealth Advisors, LLC	1,479	0.00%	$3K	+463 (+45.6%)
124	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	1,190	0.00%	$3K	NEW
125	Farther Finance Advisors, LLC	1,040	0.00%	$2K	+554 (+114.0%)
126	Rockefeller Capital Management L.P.	1,000	0.00%	$2K	—
127	SIMPLEX TRADING, LLC	932	0.00%	$2K	-389 (-29.4%)
128	ROTHSCHILD INVESTMENT LLC	559	0.00%	$1K	+256 (+84.5%)
129	Sterling Capital Management LLC	537	0.00%	$1K	-443 (-45.2%)
130	HARBOUR INVESTMENTS, INC.	501	0.00%	$1K	—
131	Essential Partners LLC	451	0.00%	$1K	-7,537 (-94.4%)
132	Rialto Wealth Management, LLC	420	0.00%	$950	—
133	N.E.W. Advisory Services LLC	325	0.00%	$735	—
134	SBI Securities Co., Ltd.	321	0.00%	$725	-9 (-2.7%)
135	Covestor Ltd	178	0.00%	$0	+126 (+242.3%)
136	True Wealth Design, LLC	178	0.00%	$403	+138 (+345.0%)
137	SJS Investment Consulting Inc.	157	0.00%	$355	+90 (+134.3%)
138	Zurcher Kantonalbank (Zurich Cantonalbank)	154	0.00%	$348	—
139	Quarry LP	123	0.00%	$278	NEW
140	Fortitude Family Office, LLC	108	0.00%	$244	NEW
141	IFP Advisors, Inc	91	0.00%	$206	+29 (+46.8%)
142	Leo H. Evart, Inc.	50	0.00%	$113	—
143	Wealth Enhancement Trust Services, Inc.	46	0.00%	$104	NEW
144	Transamerica Financial Advisors, LLC	46	0.00%	$103	—
145	Clearstead Advisors, LLC	15	0.00%	$34	NEW
146	Caitong International Asset Management Co., Ltd	12	0.00%	$27	+11 (+1100.0%)
147	JPL Wealth Management, LLC	8	0.00%	$19	NEW
148	HUNTINGTON NATIONAL BANK	2	0.00%	$5	—

Exited since 2026-03-31 (13 filers; largest 13 shown)

Timothy G. Youngquist 2020 Irrevocable Trust	−978,071
Private Advisor Group, LLC	−847,001
Diametric Capital, LP	−359,268
LAZARD ASSET MANAGEMENT LLC	−282,380
PRELUDE CAPITAL MANAGEMENT, LLC	−54,680
CWM, LLC	−39,077
StoneX Group Inc.	−14,096
Fiduciary Trust Co	−13,620
Pallas Capital Advisors LLC	−10,325
VICTORY CAPITAL MANAGEMENT INC	−10,081
MAI Capital Management	−232
Root Financial Partners, LLC	−73
NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO	−1

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting person	Form	Filed	Event	Shares	% of class
LYTTON LAURENCE W	SCHEDULE 13G/A	2026-08-13	2026-06-30	5,592,972	5.2%
Telemark Fund LP	SCHEDULE 13G	2026-07-29	2026-07-24	7,846,411	7.3%
Telemark Asset Management, LLC	SCHEDULE 13G	2026-07-29	2026-07-24	7,846,411	7.3%
Colin S. McNay	SCHEDULE 13G	2026-07-29	2026-07-24	7,846,411	7.3%
abrdn Inc.	SCHEDULE 13G/A	2026-07-20	2026-06-30	0	7.45%
Aberdeen Group plc	SCHEDULE 13G/A	2026-07-20	2026-06-30	6,769,954	7.45%
James Silverman	SCHEDULE 13G/A	2026-05-05	2026-03-31	6,785,000	7.47%
Opaleye Management Inc.	SCHEDULE 13G/A	2026-05-05	2026-03-31	6,785,000	7.47%
Opaleye, L.P.	SCHEDULE 13G/A	2026-05-05	2026-03-31	6,665,000	7.33%
Millennium Management LLC	SCHEDULE 13G	2026-04-14	2026-04-09	4,734,627	5.2%
Integrated Core Strategies (US) LLC	SCHEDULE 13G	2026-04-14	2026-04-09	4,734,627	5.2%
Israel A. Englander	SCHEDULE 13G	2026-04-14	2026-04-09	4,734,627	5.2%
Millennium Group Management LLC	SCHEDULE 13G	2026-04-14	2026-04-09	4,734,627	5.2%
The Vanguard Group	SCHEDULE 13G/A	2026-03-26	2026-03-13	0	0%
ELI CASDIN	SCHEDULE 13G/A	2025-11-14	2025-09-30	0	0%
CASDIN PARTNERS MASTER FUND, L.P.	SCHEDULE 13G/A	2025-11-14	2025-09-30	0	0%
CASDIN PARTNERS GP, LLC	SCHEDULE 13G/A	2025-11-14	2025-09-30	0	0%
CASDIN CAPITAL, LLC	SCHEDULE 13G/A	2025-11-14	2025-09-30	0	0%
FMR LLC	SCHEDULE 13G/A	2025-11-05	2025-09-30	584,367	0.6%
Abigail P. Johnson	SCHEDULE 13G/A	2025-11-05	2025-09-30	584,367	0.6%
BlackRock, Inc.	SCHEDULE 13G/A	2025-04-24	2025-03-31	5,979,480	7.2%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.