Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

Arcturus Therapeutics Holdings Inc.

CIK 0001768224ARCT· quarter ending 2026-06-30
Shares outstanding
28,423,069
Held by 13F filers
24,982,028
87.9% of shares outstanding
Holders
135
+11 vs 2026-03-31 · 20 new, 9 exited
Value
$167.9M
shares +2.9% vs prior quarter

Share classes (cover page of the 10-Q filed 2026-08-06, as of 2026-08-04)

ClassListedShares outstandingHeld by 13F filers% of classHoldersCUSIP
Common stockARCT · NASDAQ28,423,06924,982,02887.9%13503969T109

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine
CountCUSIPClass (as filers write it)HoldersShares heldClass shares% of classValueOptionsMapped via
03969T109COM13524,982,02828,423,06987.9%$167.9M2 call / 2 put13DGholders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings.

Top holders (135 of 135 · 03969T109)

change vs 2026-03-31
#FilerShares% of classValueChange
1FEDERATED HERMES, INC.3,303,64011.62%$22.2M—
2ARK Investment Management LLC2,901,63510.21%$19.5M+132,117 (+4.8%)
3BlackRock, Inc.2,614,0129.20%$17.6M+731 (+0.0%)
4Sumitomo Mitsui Trust Group, Inc.1,963,1236.91%$13.2M-325,812 (-14.2%)
5Amova Asset Management Americas, Inc.1,960,4586.90%$13.1M-328,477 (-14.4%)
6VANGUARD CAPITAL MANAGEMENT LLC1,114,9563.92%$7.5M+7,936 (+0.7%)
7TWO SIGMA INVESTMENTS, LP1,091,1373.84%$7.3M+9,353 (+0.9%)
8VANGUARD PORTFOLIO MANAGEMENT LLC951,5753.35%$6.4M-26,347 (-2.7%)
9GEODE CAPITAL MANAGEMENT, LLC693,6602.44%$4.7M+32,732 (+5.0%)
10AQR CAPITAL MANAGEMENT LLC675,0142.37%$4.5M+513,736 (+318.5%)
11JACOBS LEVY EQUITY MANAGEMENT, INC647,5112.28%$4.4M+448,783 (+225.8%)
12STATE STREET CORP569,5982.00%$3.8M+25,735 (+4.7%)
13UBS Group AG490,6321.73%$3.3M+32,355 (+7.1%)
14D. E. Shaw & Co., Inc.392,3581.38%$2.6M+140,324 (+55.7%)
15Empire Life Investments Inc.366,2171.29%$2.5M-17,276 (-4.5%)
16Hennion & Walsh Asset Management, Inc.346,2071.22%$2.3M+66,528 (+23.8%)
17MILLENNIUM MANAGEMENT LLC336,9961.19%$2.3M-141,561 (-29.6%)
18GOLDMAN SACHS GROUP INC309,8631.09%$2.1M+111,182 (+56.0%)
19BNP PARIBAS FINANCIAL MARKETS261,7890.92%$1.8M+66,929 (+34.3%)
20CHARLES SCHWAB INVESTMENT MANAGEMENT INC231,6280.81%$1.6M+15,128 (+7.0%)
21Point72 Asset Management, L.P.231,5820.81%$1.6M+167,737 (+262.7%)
22DIMENSIONAL FUND ADVISORS LP221,3220.78%$1.5M-67,033 (-23.2%)
23HEARTLAND ADVISORS INC219,3000.77%$1.5M-36,700 (-14.3%)
24NORTHERN TRUST CORP204,5780.72%$1.4M-15,035 (-6.8%)
25JPMORGAN CHASE & CO195,7040.69%$1.3M-2,111 (-1.1%)
26VANGUARD FIDUCIARY TRUST CO174,0480.61%$1.2M+4,236 (+2.5%)
27Jump Financial, LLC138,2090.49%$929K+69,916 (+102.4%)
28ACADIAN ASSET MANAGEMENT LLC131,4510.46%$880K-149,049 (-53.1%)
29IQ EQ FUND MANAGEMENT (IRELAND) Ltd126,3900.44%$849K+12,135 (+10.6%)
30Invesco Ltd.104,1880.37%$700K-9,382 (-8.3%)
31MORGAN STANLEY97,4060.34%$655K-28,729 (-22.8%)
32Quantinno Capital Management LP85,8750.30%$577K+14,522 (+20.4%)
33Bank of New York Mellon Corp80,3140.28%$540K+5,317 (+7.1%)
34MARSHALL WACE, LLP77,2690.27%$519KNEW
35Russell Investments Group, Ltd.75,9030.27%$510K+44,199 (+139.4%)
36AMERICAN CENTURY COMPANIES INC65,3610.23%$439K-651 (-1.0%)
37AMERIPRISE FINANCIAL INC65,2820.23%$439KNEW
38MASSACHUSETTS FINANCIAL SERVICES CO /MA/57,4800.20%$386K-2,290 (-3.8%)
39HSBC HOLDINGS PLC56,8950.20%$394K+4,070 (+7.7%)
40Nuveen, LLC52,5420.18%$353K—
41Squarepoint Ops LLC50,2530.18%$338K+36,479 (+264.8%)
42GSA CAPITAL PARTNERS LLP49,1450.17%$330K-84,565 (-63.2%)
43SUSQUEHANNA INTERNATIONAL GROUP, LLP47,5570.17%$320K-156,457 (-76.7%)
44Y-Intercept (Hong Kong) Ltd43,2910.15%$291KNEW
45CITADEL ADVISORS LLC40,9460.14%$275K+6,164 (+17.7%)
46nVerses Capital, LLC38,9500.14%$262K+714 (+1.9%)
47PRICE T ROWE ASSOCIATES INC /MD/36,8870.13%$248K+7,094 (+23.8%)
48BRIDGEWAY CAPITAL MANAGEMENT, LLC36,5000.13%$245K—
49ALLIANCEBERNSTEIN L.P.35,2800.12%$272K+7,560 (+27.3%)
50PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO34,0500.12%$229K+26,399 (+345.0%)
51RHUMBLINE ADVISERS31,8870.11%$214K+1,011 (+3.3%)
52WELLS FARGO & COMPANY/MN29,8920.11%$201K+10,888 (+57.3%)
53XTX Topco Ltd29,2820.10%$197KNEW
54DEUTSCHE BANK AG\28,9940.10%$195K-2,370 (-7.6%)
55ALPS ADVISORS INC28,8450.10%$194K+5,705 (+24.7%)
56Engineers Gate Manager LP28,7680.10%$193K-17,065 (-37.2%)
57Brevan Howard Capital Management LP28,7620.10%$193K-3,982 (-12.2%)
58TritonPoint Partners, LLC28,1860.10%$189K+114 (+0.4%)
59Allspring Global Investments Holdings, LLC27,6510.10%$186K+781 (+2.9%)
60Green Alpha Advisors, LLC27,0910.10%$182K-1,047 (-3.7%)
61Schonfeld Strategic Advisors LLC26,3150.09%$177KNEW
62MGO ONE SEVEN LLC24,7880.09%$167K-2,029 (-7.6%)
63MERCER GLOBAL ADVISORS INC /ADV24,0060.08%$161K+13,118 (+120.5%)
64Apollon Wealth Management, LLC23,4330.08%$157KNEW
65BANK OF AMERICA CORP /DE/22,5010.08%$151K-6,250 (-21.7%)
66RENAISSANCE TECHNOLOGIES LLC21,4000.08%$144K-156,900 (-88.0%)
67Mariner, LLC20,5550.07%$138KNEW
68FMR LLC19,5420.07%$131K+14,921 (+322.9%)
69Walleye Capital LLC19,3230.07%$130KNEW
70Freestone Capital Holdings, LLC18,6550.07%$125K—
71AXQ CAPITAL, LP18,2770.06%$123K-4,705 (-20.5%)
72COMMONWEALTH EQUITY SERVICES, LLC17,7410.06%$119K-426 (-2.3%)
73Corient Private Wealth LP17,3540.06%$117KNEW
74CANADA LIFE ASSURANCE Co17,3530.06%$116K+4,559 (+35.6%)
75Abel Hall, LLC16,1420.06%$108KNEW
76Diversify Advisory Services, LLC15,7850.06%$91KNEW
77DIVERSIFY WEALTH MANAGEMENT, LLC15,7850.06%$91K—
78Susquehanna Portfolio Strategies, LLC15,7300.06%$106K-48,666 (-75.6%)
79MetLife Investment Management, LLC15,3660.05%$103K+8,541 (+125.1%)
80Dynamic Technology Lab Private Ltd15,1770.05%$102KNEW
81JANE STREET GROUP, LLC15,0100.05%$101K-8,073 (-35.0%)
82RAYMOND JAMES FINANCIAL INC14,3230.05%$96KNEW
83BARCLAYS PLC13,7750.05%$93K+145 (+1.1%)
84WEALTH ENHANCEMENT ADVISORY SERVICES, LLC13,2820.05%$93KNEW
85Quadrature Capital Ltd13,0750.05%$88K-17,003 (-56.5%)
86CITIGROUP INC13,0270.05%$88K+10,700 (+459.8%)
87MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.12,3410.04%$83K-21,961 (-64.0%)
88TWO SIGMA SECURITIES, LLC11,8640.04%$80KNEW
89Inscription Capital, LLC11,7740.04%$79K—
90Arax Advisory Partners11,7110.04%$79K+11,649 (+18788.7%)
91Formidable Asset Management, LLC11,7000.04%$90K+1,100 (+10.4%)
92Jain Global LLC10,0850.04%$68K-11,103 (-52.4%)
93Vanguard Global Advisers, LLC8,5070.03%$57K—
94NEW YORK STATE COMMON RETIREMENT FUND8,4970.03%$57K—
95Police & Firemen's Retirement System of New Jersey6,9570.02%$47K—
96China Universal Asset Management Co., Ltd.4,5240.02%$30K-518 (-10.3%)
97NISA INVESTMENT ADVISORS, LLC3,3000.01%$22KNEW
98Ameritas Investment Partners, Inc.2,3170.01%$16K—
99Legal & General Group Plc2,3040.01%$15K—
100SIGNATUREFD, LLC2,1880.01%$15K—
101UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC1,7870.01%$12K—
102GAMMA Investing LLC1,7500.01%$12K+244 (+16.2%)
103ROYAL BANK OF CANADA1,6370.01%$11K+670 (+69.3%)
104Tower Research Capital LLC (TRC)1,5500.01%$10K-2,873 (-65.0%)
105OSAIC HOLDINGS, INC.1,4510.01%$10K-949 (-39.5%)
106Mirae Asset Global Investments Co., Ltd.1,2710.00%$9K-211 (-14.2%)
107PNC Financial Services Group, Inc.1,0450.00%$7K-13 (-1.2%)
108SBI Securities Co., Ltd.9210.00%$6K+524 (+132.0%)
109Essential Partners LLC8240.00%$6K+488 (+145.2%)
110Bank Julius Baer & Co. Ltd, Zurich8000.00%$5K—
111Allworth Financial LP7890.00%$5K+593 (+302.6%)
112Global Retirement Partners, LLC6720.00%$5KNEW
113GF FUND MANAGEMENT CO. LTD.6110.00%$4K-42 (-6.4%)
114SIMPLEX TRADING, LLC6070.00%$4K-17,559 (-96.7%)
115IFP Advisors, Inc5840.00%$4K+555 (+1913.8%)
116Farther Finance Advisors, LLC5550.00%$4K+226 (+68.7%)
117Sterling Capital Management LLC5400.00%$4K-71 (-11.6%)
118EverSource Wealth Advisors, LLC4360.00%$3K+325 (+292.8%)
119US BANCORP \DE\4230.00%$3K—
120Versant Capital Management, Inc4050.00%$3K+90 (+28.6%)
121CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM3690.00%$2K-1,001 (-73.1%)
122ROTHSCHILD INVESTMENT LLC3230.00%$2K+256 (+382.1%)
123LOS ANGELES CAPITAL MANAGEMENT LLC2570.00%$2K—
124True Wealth Design, LLC2530.00%$2K+192 (+314.8%)
125Federation des caisses Desjardins du Quebec2350.00%$2K—
126SJS Investment Consulting Inc.1720.00%$1K+170 (+8500.0%)
127Fortitude Family Office, LLC1340.00%$900NEW
128Steward Partners Investment Advisory, LLC1050.00%$706+3 (+2.9%)
129Optiver Holding B.V.920.00%$618—
130Parkside Financial Bank & Trust800.00%$538NEW
131Caitong International Asset Management Co., Ltd680.00%$457-955 (-93.4%)
132Clearstead Advisors, LLC550.00%$370NEW
133Covestor Ltd420.00%$0+30 (+250.0%)
134NFSG Corp400.00%$269—
135HARBOUR INVESTMENTS, INC.310.00%$211+18 (+138.5%)
Exited since 2026-03-31 (9 filers; largest 9 shown)
Odyssean, LLC−22,956
SG Americas Securities, LLC−20,972
Castleview Partners, LLC−16,857
HRT FINANCIAL LP−15,129
ADAR1 Capital Management, LLC−15,000
CWM, LLC−9,324
MAI Capital Management−465
Wealth Preservation Advisors, LLC−40
Root Financial Partners, LLC−22

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting personFormFiledEventShares% of class
ARK Investment Management LLCSCHEDULE 13G/A2026-05-072026-04-302,947,55610.37%
Catherine D. WoodSCHEDULE 13G/A2026-05-072026-04-302,947,55610.37%
The Vanguard GroupSCHEDULE 13G/A2026-03-262026-03-1300%
Dmitry BalyasnySCHEDULE 13G/A2026-02-172025-12-3100%
Dames GP LLCSCHEDULE 13G/A2026-02-172025-12-3100%
Balyasny Asset Management L.P.SCHEDULE 13G/A2026-02-172025-12-3100%
BAM GP LLCSCHEDULE 13G/A2026-02-172025-12-3100%
Balyasny Asset Management Holdings LPSCHEDULE 13G/A2026-02-172025-12-3100%
J. Christopher DonahueSCHEDULE 13G/A2026-01-082025-12-313,303,64011.63%
Ann C. DonahueSCHEDULE 13G/A2026-01-082025-12-313,303,64011.63%
Federated Hermes, Inc.SCHEDULE 13G/A2026-01-082025-12-313,303,64011.63%
Thomas R. DonahueSCHEDULE 13G/A2026-01-082025-12-313,303,64011.63%
Voting Shares Irrevocable TrustSCHEDULE 13G/A2026-01-082025-12-313,303,64011.63%
Amova Asset Management Americas, Inc.SCHEDULE 13G/A2025-11-132025-09-301,947,8357.2%
Sumitomo Mitsui Trust Group, Inc.SCHEDULE 13G/A2025-10-272025-09-301,947,8357.2%
Amova Asset Management Co., Ltd.SCHEDULE 13G/A2025-10-272025-09-301,947,8357.2%
Nikko Asset Management Americas, Inc.SCHEDULE 13G/A2025-05-122025-03-311,926,0847.1%
Nikko Asset Management Co., Ltd.SCHEDULE 13G/A2025-04-252025-03-311,926,0847.1%
BlackRock, Inc.SCHEDULE 13G/A2025-04-232025-03-312,104,5377.8%
STATE STREET CORPORATIONSCHEDULE 13G2025-02-052024-12-311,326,1584.9%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.