Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

Denali Therapeutics Inc.

CIK 0001714899DNLI· quarter ending 2026-06-30

Shares outstanding

159,847,403

as of 2026-07-31 · 10-Q cover page, filed 2026-08-06

Held by 13F filers

156,230,244

97.7% of shares outstanding

Holders

294

+25 vs 2026-03-31 · 56 new, 31 exited

Value

$4.5B

shares +5.7% vs prior quarter

Share classes (cover page of the 10-Q filed 2026-08-06, as of 2026-07-31)

Class	Listed	Shares outstanding	Held by 13F filers	% of class	Holders	CUSIP
Common stock	DNLI · NASDAQ	159,847,403	156,230,244	97.7%	294	24823R105

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine

Count	CUSIP	Class (as filers write it)	Holders	Shares held	Class shares	% of class	Value	Options	Mapped via
	24823R105	COM	294	156,230,244	159,847,403	97.7%	$4.5B	4 call / 4 put	13DG	holders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings.

Top holders (250 of 294 · 24823R105)

change vs 2026-03-31

#	Filer	Shares	% of class	Value	Change
1	T. Rowe Price Investment Management, Inc.	16,178,963	10.12%	$416.1M	+7,659,771 (+89.9%)
2	BlackRock, Inc.	14,026,413	8.77%	$360.8M	+991,979 (+7.6%)
3	BAILLIE GIFFORD & CO	11,960,230	7.48%	$307.6M	-106,249 (-0.9%)
4	STATE STREET CORP	7,049,304	4.41%	$181.3M	+777,303 (+12.4%)
5	Temasek Holdings (Private) Ltd	7,012,974	4.39%	$180.4M	—
6	VANGUARD PORTFOLIO MANAGEMENT LLC	6,477,057	4.05%	$166.6M	+356,897 (+5.8%)
7	VANGUARD CAPITAL MANAGEMENT LLC	6,275,729	3.93%	$161.4M	+127,772 (+2.1%)
8	FMR LLC	4,535,836	2.84%	$116.7M	-39,538 (-0.9%)
9	GOLDMAN SACHS GROUP INC	4,155,895	2.60%	$106.9M	-251,360 (-5.7%)
10	GEODE CAPITAL MANAGEMENT, LLC	3,973,342	2.49%	$102.2M	+246,228 (+6.6%)
11	Holocene Advisors, LP	3,905,225	2.44%	$100.4M	+709,796 (+22.2%)
12	BAKER BROS. ADVISORS LP	3,734,336	2.34%	$96.0M	+2,641 (+0.1%)
13	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	3,161,972	1.98%	$81.3M	-1,270,342 (-28.7%)
14	PRICE T ROWE ASSOCIATES INC /MD/	3,136,149	1.96%	$80.7M	+672,003 (+27.3%)
15	UBS Group AG	3,120,592	1.95%	$80.3M	+549,248 (+21.4%)
16	ALLIANCEBERNSTEIN L.P.	2,984,807	1.87%	$57.3M	+91,189 (+3.2%)
17	DIMENSIONAL FUND ADVISORS LP	2,860,604	1.79%	$73.6M	+168,429 (+6.3%)
18	NORGES BANK	2,824,746	1.77%	$72.7M	NEW
19	Pictet Asset Management Holding SA	2,791,206	1.75%	$71.9M	+232,341 (+9.1%)
20	Capital Research Global Investors	2,565,817	1.61%	$66.0M	+51,229 (+2.0%)
21	Fisher Asset Management, LLC	2,414,162	1.51%	$62.1M	-22,733 (-0.9%)
22	NOMURA ASSET MANAGEMENT INTERNATIONAL INC.	2,208,263	1.38%	$56.8M	+2,093,739 (+1828.2%)
23	Casdin Capital, LLC	2,045,000	1.28%	$52.6M	+205,000 (+11.1%)
24	Crestline Management, L.P.	1,910,732	1.20%	$49.1M	-300,000 (-13.6%)
25	PRINCIPAL FINANCIAL GROUP INC	1,420,083	0.89%	$36.5M	-58,403 (-4.0%)
26	Foresite Capital Management VI LLC	1,396,771	0.87%	$35.9M	+186,800 (+15.4%)
27	Siren, L.L.C.	1,396,513	0.87%	$35.9M	—
28	NORTHERN TRUST CORP	1,389,765	0.87%	$35.7M	+79,109 (+6.0%)
29	BNP PARIBAS ASSET MANAGEMENT Holding S.A.	1,355,514	0.85%	$34.9M	+1,059,011 (+357.2%)
30	Aberdeen Group plc	1,345,481	0.84%	$34.6M	-508,965 (-27.4%)
31	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	1,244,955	0.78%	$32.0M	+37,556 (+3.1%)
32	BANK OF AMERICA CORP /DE/	1,230,957	0.77%	$31.7M	+96,774 (+8.5%)
33	Artisan Partners Limited Partnership	1,224,497	0.77%	$31.5M	+124,778 (+11.3%)
34	Candriam S.C.A.	1,189,886	0.74%	$30.6M	+90,041 (+8.2%)
35	MILLENNIUM MANAGEMENT LLC	1,125,748	0.70%	$29.0M	-132,202 (-10.5%)
36	ADAR1 Capital Management, LLC	1,029,080	0.64%	$26.5M	-842,375 (-45.0%)
37	VANGUARD FIDUCIARY TRUST CO	991,904	0.62%	$25.5M	+42,209 (+4.4%)
38	MORGAN STANLEY	961,050	0.60%	$24.7M	-143,707 (-13.0%)
39	Bank of New York Mellon Corp	800,283	0.50%	$20.6M	+32,244 (+4.2%)
40	DAFNA Capital Management LLC	739,014	0.46%	$19.0M	+420,514 (+132.0%)
41	AMERICAN CENTURY COMPANIES INC	712,526	0.45%	$18.3M	-223,166 (-23.9%)
42	Cercano Management LLC	625,000	0.39%	$16.1M	—
43	Nuveen, LLC	608,967	0.38%	$15.7M	+313,381 (+106.0%)
44	Ensign Peak Advisors, Inc	582,669	0.36%	$15.0M	+66,948 (+13.0%)
45	MARSHALL WACE, LLP	549,830	0.34%	$14.1M	-441,438 (-44.5%)
46	SEGALL BRYANT & HAMILL, LLC	519,483	0.32%	$13.4M	+8,940 (+1.8%)
47	CITIGROUP INC	487,278	0.30%	$12.5M	+448,000 (+1140.6%)
48	Defilade Capital Management, L.P.	391,866	0.25%	$10.1M	-968,103 (-71.2%)
49	AQR CAPITAL MANAGEMENT LLC	388,469	0.24%	$10.0M	+285,890 (+278.7%)
50	Woodline Partners LP	364,082	0.23%	$9.4M	—
51	Invesco Ltd.	343,898	0.22%	$8.8M	+33,702 (+10.9%)
52	ExodusPoint Capital Management, LP	336,567	0.21%	$8.7M	NEW
53	Marex Group Ltd	295,631	0.18%	$7.6M	+8,631 (+3.0%)
54	JPMORGAN CHASE & CO	287,019	0.18%	$7.6M	-36,970 (-11.4%)
55	Swiss National Bank	270,500	0.17%	$7.0M	-3,300 (-1.2%)
56	TD Asset Management Inc	231,644	0.14%	$6.0M	-24,122 (-9.4%)
57	GROUPAMA ASSET MANAGMENT	229,939	0.14%	$5.9M	+22,809 (+11.0%)
58	Privium Fund Management B.V.	229,857	0.14%	$5.9M	-25,000 (-9.8%)
59	XTX Topco Ltd	226,953	0.14%	$5.8M	+148,386 (+188.9%)
60	Alyeska Investment Group, L.P.	219,334	0.14%	$5.6M	-1,721,789 (-88.7%)
61	Susquehanna Portfolio Strategies, LLC	211,474	0.13%	$5.4M	+53,578 (+33.9%)
62	MASTERS CAPITAL MANAGEMENT LLC	200,000	0.13%	$5.1M	-400,000 (-66.7%)
63	Burkehill Global Management, LP	200,000	0.13%	$5.1M	—
64	SCS Capital Management LLC	197,998	0.12%	$5.1M	—
65	RHUMBLINE ADVISERS	194,787	0.12%	$5.0M	+10,539 (+5.7%)
66	WELLS FARGO & COMPANY/MN	189,865	0.12%	$4.9M	+82,208 (+76.4%)
67	Universal- Beteiligungs- und Servicegesellschaft mbH	179,195	0.11%	$4.6M	+79,692 (+80.1%)
68	Rhenman & Partners Asset Management AB	179,000	0.11%	$4.6M	-20,000 (-10.1%)
69	NEW YORK STATE COMMON RETIREMENT FUND	171,712	0.11%	$4.4M	+43,593 (+34.0%)
70	TEACHERS RETIREMENT SYSTEM OF THE STATE OF KENTUCKY	166,181	0.10%	$4.3M	—
71	EDMOND DE ROTHSCHILD HOLDING S.A.	165,080	0.10%	$4.2M	-60,600 (-26.9%)
72	ALPS ADVISORS INC	160,378	0.10%	$4.1M	+31,273 (+24.2%)
73	E. Ohman J:or Asset Management AB	160,000	0.10%	$4.1M	NEW
74	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	152,086	0.10%	$3.9M	-10,490 (-6.5%)
75	DEUTSCHE BANK AG\	149,804	0.09%	$3.9M	+29,436 (+24.5%)
76	GREAT LAKES ADVISORS, LLC	145,045	0.09%	$3.7M	-82,065 (-36.1%)
77	VANGUARD ASSET MANAGEMENT, Ltd	141,153	0.09%	$3.6M	+15,910 (+12.7%)
78	Man Group plc	139,065	0.09%	$3.6M	+20 (+0.0%)
79	Hudson Bay Capital Management LP	130,000	0.08%	$3.3M	-216,360 (-62.5%)
80	Odyssean, LLC	125,062	0.08%	$3.2M	NEW
81	SUSQUEHANNA INTERNATIONAL GROUP, LLP	123,258	0.08%	$3.2M	+72,783 (+144.2%)
82	MANUFACTURERS LIFE INSURANCE COMPANY, THE	117,348	0.07%	$3.0M	-8,094 (-6.5%)
83	Zurcher Kantonalbank (Zurich Cantonalbank)	115,775	0.07%	$3.0M	+4,081 (+3.7%)
84	B. Metzler seel. Sohn & Co. AG	112,300	0.07%	$2.9M	—
85	Legal & General Group Plc	111,953	0.07%	$2.9M	+1,703 (+1.5%)
86	Integrated Wealth Concepts LLC	102,241	0.06%	$2.6M	+2,687 (+2.7%)
87	PFM Health Sciences, LP	100,966	0.06%	$2.6M	NEW
88	Ghisallo Capital Management LLC	100,000	0.06%	$2.6M	—
89	Integral Health Asset Management, LLC	100,000	0.06%	$2.6M	NEW
90	Lumbard & Kellner, LLC	98,282	0.06%	$2.5M	+3,500 (+3.7%)
91	Rafferty Asset Management, LLC	96,342	0.06%	$2.5M	-54,345 (-36.1%)
92	D.A. DAVIDSON & CO.	87,741	0.05%	$2.3M	-22,494 (-20.4%)
93	Russell Investments Group, Ltd.	86,032	0.05%	$2.2M	+71,974 (+512.0%)
94	MetLife Investment Management, LLC	85,621	0.05%	$2.2M	+8,063 (+10.4%)
95	BARCLAYS PLC	84,892	0.05%	$2.2M	+32,808 (+63.0%)
96	STATE OF WISCONSIN INVESTMENT BOARD	84,652	0.05%	$2.2M	+2,295 (+2.8%)
97	Creative Planning	84,644	0.05%	$2.2M	+10,575 (+14.3%)
98	AMERIPRISE FINANCIAL INC	82,650	0.05%	$2.1M	-6,884 (-7.7%)
99	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	78,937	0.05%	$2.0M	-16,256 (-17.1%)
100	Phocas Financial Corp.	71,197	0.04%	$1.8M	NEW
101	Knights of Columbus Asset Advisors LLC	67,591	0.04%	$1.7M	+11,262 (+20.0%)
102	CITADEL ADVISORS LLC	66,971	0.04%	$1.7M	-114,446 (-63.1%)
103	1492 Capital Management LLC	66,368	0.04%	$1.7M	-1,049 (-1.6%)
104	Vanguard Global Advisers, LLC	61,400	0.04%	$1.6M	+2,621 (+4.5%)
105	Jupiter Topco LLC	60,028	0.04%	$1.5M	NEW
106	Dana Investment Advisors, Inc.	59,971	0.04%	$1.5M	+8,049 (+15.5%)
107	DekaBank Deutsche Girozentrale	59,800	0.04%	$1.1M	—
108	Handelsbanken Fonder AB	59,600	0.04%	$1.5M	+15,800 (+36.1%)
109	SEI INVESTMENTS CO	58,989	0.04%	$1.5M	+6,754 (+12.9%)
110	PRUDENTIAL FINANCIAL INC	58,135	0.04%	$1.5M	+27,565 (+90.2%)
111	Engineers Gate Manager LP	57,430	0.04%	$1.5M	+41,320 (+256.5%)
112	FRANKLIN RESOURCES INC	56,662	0.04%	$1.5M	+7,930 (+16.3%)
113	Pathstone Holdings, LLC	54,014	0.03%	$1.4M	-2,311 (-4.1%)
114	Virtus Investment Advisers, LLC	53,688	0.03%	$1.4M	+38,522 (+254.0%)
115	WELLINGTON MANAGEMENT GROUP LLP	52,618	0.03%	$1.4M	-745 (-1.4%)
116	IRONWOOD INVESTMENT MANAGEMENT LLC	52,080	0.03%	$1.3M	+18 (+0.0%)
117	ENVESTNET ASSET MANAGEMENT INC	51,424	0.03%	$1.3M	+524 (+1.0%)
118	PROFUND ADVISORS LLC	49,899	0.03%	$1.3M	-18,328 (-26.9%)
119	MAI Capital Management	49,160	0.03%	$1.3M	+2,570 (+5.5%)
120	VOYA INVESTMENT MANAGEMENT LLC	45,823	0.03%	$1.2M	+22,151 (+93.6%)
121	Savvy Advisors, Inc.	45,194	0.03%	$1.2M	NEW
122	Employees Retirement System of Texas	44,340	0.03%	$1.1M	-2,837 (-6.0%)
123	RAYMOND JAMES FINANCIAL INC	43,744	0.03%	$1.1M	+9,361 (+27.2%)
124	State of New Jersey Common Pension Fund D	43,587	0.03%	$1.1M	-3,884 (-8.2%)
125	Police & Firemen's Retirement System of New Jersey	43,387	0.03%	$1.1M	+4,842 (+12.6%)
126	SG Americas Securities, LLC	42,339	0.03%	$1.1M	-4,303 (-9.2%)
127	TUDOR INVESTMENT CORP ET AL	40,965	0.03%	$1.1M	NEW
128	Jain Global LLC	38,435	0.02%	$989K	NEW
129	Quantinno Capital Management LP	37,651	0.02%	$968K	+7,481 (+24.8%)
130	Vanguard Investments Australia, Ltd.	37,410	0.02%	$962K	—
131	ARIZONA STATE RETIREMENT SYSTEM	37,196	0.02%	$957K	-957 (-2.5%)
132	NOMURA ASSET MANAGEMENT CO LTD	36,732	0.02%	$945K	—
133	INTECH INVESTMENT MANAGEMENT LLC	35,484	0.02%	$913K	-3,447 (-8.9%)
134	Jefferies Financial Group Inc.	34,425	0.02%	$885K	—
135	STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM	34,168	0.02%	$879K	—
136	Mariner, LLC	32,799	0.02%	$844K	+14,668 (+80.9%)
137	Thrivent Financial for Lutherans	32,555	0.02%	$837K	—
138	Yorktown Management & Research Co Inc	32,100	0.02%	$826K	NEW
139	J. Safra Sarasin Holding AG	31,250	0.02%	$804K	NEW
140	KLP KAPITALFORVALTNING AS	30,900	0.02%	$795K	—
141	China Universal Asset Management Co., Ltd.	29,819	0.02%	$767K	-10,824 (-26.6%)
142	SBI Okasan Asset Management Co.Ltd.	29,500	0.02%	$764K	—
143	Erste Asset Management GmbH	26,900	0.02%	$696K	—
144	BNP PARIBAS FINANCIAL MARKETS	25,071	0.02%	$645K	-14,822 (-37.2%)
145	ProShare Advisors LLC	24,466	0.02%	$629K	-3,271 (-11.8%)
146	SIMPLEX TRADING, LLC	23,952	0.01%	$616K	+11,637 (+94.5%)
147	Cerity Partners LLC	21,587	0.01%	$555K	+8,929 (+70.5%)
148	CAPSTONE INVESTMENT ADVISORS, LLC	20,964	0.01%	$539K	+8,001 (+61.7%)
149	IEQ CAPITAL, LLC	20,185	0.01%	$519K	NEW
150	ALLSTATE CORP	19,313	0.01%	$497K	NEW
151	Connor, Clark & Lunn Investment Management Ltd.	18,556	0.01%	$477K	NEW
152	MARTINGALE ASSET MANAGEMENT L P	18,530	0.01%	$477K	-5 (-0.0%)
153	Banque Cantonale Vaudoise	17,926	0.01%	$461.0M	NEW
154	Fieldview Capital Management, LLC	17,873	0.01%	$460K	NEW
155	State of Wyoming	17,778	0.01%	$457K	NEW
156	Bellevue Group AG	17,766	0.01%	$457K	—
157	VICTORY CAPITAL MANAGEMENT INC	17,197	0.01%	$442K	+1,003 (+6.2%)
158	OSAIC HOLDINGS, INC.	17,016	0.01%	$438K	+603 (+3.7%)
159	INTRUST BANK NA	16,477	0.01%	$424K	+2,138 (+14.9%)
160	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	16,467	0.01%	$423K	NEW
161	MERCER GLOBAL ADVISORS INC /ADV	16,257	0.01%	$418K	NEW
162	HIMENSION CAPITAL (SINGAPORE) PTE. LTD.	15,949	0.01%	$410K	NEW
163	TEACHER RETIREMENT SYSTEM OF TEXAS	15,713	0.01%	$404K	-2,169 (-12.1%)
164	Graham Capital Management, L.P.	15,669	0.01%	$403K	NEW
165	Breakwater Capital Group	15,601	0.01%	$401K	+15 (+0.1%)
166	CONTRAVISORY INVESTMENT MANAGEMENT, INC.	14,674	0.01%	$377K	+223 (+1.5%)
167	XPONANCE LLC	14,616	0.01%	$376K	+1,376 (+10.4%)
168	Empowered Funds, LLC	14,428	0.01%	$371K	-1,359 (-8.6%)
169	VIRGINIA RETIREMENT SYSTEMS ET Al	14,300	0.01%	$368K	NEW
170	WINTON GROUP Ltd	14,100	0.01%	$363K	-57,079 (-80.2%)
171	ROYAL BANK OF CANADA	13,974	0.01%	$359K	+6,439 (+85.5%)
172	LPL Financial LLC	13,868	0.01%	$357K	+1,594 (+13.0%)
173	GUGGENHEIM CAPITAL LLC	13,636	0.01%	$351K	-189 (-1.4%)
174	INCEPTIONR LLC	13,632	0.01%	$351K	NEW
175	E Fund Management Co., Ltd.	13,585	0.01%	$349K	-5,357 (-28.3%)
176	CANADA LIFE ASSURANCE Co	13,434	0.01%	$345K	—
177	Ameritas Investment Partners, Inc.	13,351	0.01%	$343K	+1,697 (+14.6%)
178	B. Riley Wealth Advisors, Inc.	12,813	0.01%	$332K	+600 (+4.9%)
179	PIER 88 INVESTMENT PARTNERS LLC	12,700	0.01%	$327K	-52,100 (-80.4%)
180	PUBLIC EMPLOYEES RETIREMENT ASSOCIATION OF COLORADO	12,238	0.01%	$315K	—
181	Atria Investments, Inc	12,130	0.01%	$312K	+18 (+0.1%)
182	Vanguard Personalized Indexing Management, LLC	11,953	0.01%	$307K	+331 (+2.8%)
183	Corient Private Wealth LP	11,704	0.01%	$301K	NEW
184	Lynx1 Capital Management LP	11,658	0.01%	$300K	-1,868,645 (-99.4%)
185	Pinnacle Wealth Planning Services, Inc.	11,587	0.01%	$298K	+181 (+1.6%)
186	Cetera Investment Advisers	11,569	0.01%	$298K	+510 (+4.6%)
187	Bayforest Capital Ltd	11,437	0.01%	$294K	+11,370 (+16970.1%)
188	Tower Research Capital LLC (TRC)	11,393	0.01%	$293K	-9,585 (-45.7%)
189	Gilbert & Cook, Inc.	11,179	0.01%	$288K	+308 (+2.8%)
190	HighTower Advisors, LLC	11,020	0.01%	$283K	-951 (-7.9%)
191	NEUBERGER BERMAN GROUP LLC	9,742	0.01%	$251K	NEW
192	PNC Financial Services Group, Inc.	9,122	0.01%	$235K	-98 (-1.1%)
193	Swiss Life Asset Management Ltd	9,091	0.01%	$234K	NEW
194	Rockefeller Capital Management L.P.	8,894	0.01%	$229K	-17,041 (-65.7%)
195	DNB Asset Management AS	8,581	0.01%	$221K	-2,378 (-21.7%)
196	Mitsubishi UFJ Trust & Banking Corp	8,494	0.01%	$218K	NEW
197	PSP Research LLC	8,463	0.01%	$218K	NEW
198	ELEVATION POINT WEALTH PARTNERS, LLC	8,343	0.01%	$215K	NEW
199	MONECO ADVISORS, LLC	8,278	0.01%	$213K	NEW
200	CIBC Bancorp USA Inc.	8,122	0.01%	$209K	NEW
201	LETKO, BROSSEAU & ASSOCIATES INC	7,850	0.00%	$202K	NEW
202	FOX RUN MANAGEMENT, L.L.C.	7,779	0.00%	$200K	NEW
203	Mirae Asset Global Investments Co., Ltd.	7,098	0.00%	$183K	-1,174 (-14.2%)
204	ACADIAN ASSET MANAGEMENT LLC	6,476	0.00%	$166K	NEW
205	ASSETMARK, INC	5,216	0.00%	$134K	+17 (+0.3%)
206	Eurizon Capital SGR S.p.A.	5,045	0.00%	$131K	—
207	KBC Group NV	4,943	0.00%	$127K	—
208	STRS OHIO	4,300	0.00%	$111K	NEW
209	NISA INVESTMENT ADVISORS, LLC	3,697	0.00%	$95K	+1,273 (+52.5%)
210	GF FUND MANAGEMENT CO. LTD.	3,357	0.00%	$86K	+63 (+1.9%)
211	FIFTH THIRD BANCORP	3,019	0.00%	$78K	+172 (+6.0%)
212	Sterling Capital Management LLC	2,837	0.00%	$73K	-1,656 (-36.9%)
213	Graticule Asia Macro Advisors LLC	2,206	0.00%	$57K	NEW
214	Allworth Financial LP	1,834	0.00%	$47K	+1,554 (+555.0%)
215	Arax Advisory Partners	1,661	0.00%	$43K	+1,360 (+451.8%)
216	Versant Capital Management, Inc	1,633	0.00%	$42K	+157 (+10.6%)
217	Pacer Advisors, Inc.	1,609	0.00%	$41K	NEW
218	GAMMA Investing LLC	1,549	0.00%	$40K	+121 (+8.5%)
219	BARTLETT & CO. WEALTH MANAGEMENT LLC	1,500	0.00%	$39K	—
220	Nykredit A/S	1,488	0.00%	$38K	NEW
221	SBI Securities Co., Ltd.	1,448	0.00%	$37K	+1,080 (+293.5%)
222	Kemnay Advisory Services Inc.	1,304	0.00%	$34K	+170 (+15.0%)
223	Smartleaf Asset Management LLC	1,107	0.00%	$29K	—
224	Global Retirement Partners, LLC	1,103	0.00%	$28K	NEW
225	Mitsubishi UFJ Asset Management Co., Ltd.	1,001	0.00%	$26K	—
226	US BANCORP \DE\	950	0.00%	$24K	+48 (+5.3%)
227	LANARK FINANCIAL, INC.	938	0.00%	$24K	-6,679 (-87.7%)
228	Optiver Holding B.V.	776	0.00%	$20K	-90 (-10.4%)
229	GROUP ONE TRADING LLC	776	0.00%	$20K	NEW
230	Nemes Rush Group LLC	700	0.00%	$18K	—
231	LOS ANGELES CAPITAL MANAGEMENT LLC	627	0.00%	$16K	-300 (-32.4%)
232	UMB Bank, n.a.	624	0.00%	$16K	—
233	Mowery & Schoenfeld Wealth Management, LLC	555	0.00%	$14K	+55 (+11.0%)
234	Farther Finance Advisors, LLC	537	0.00%	$14K	+282 (+110.6%)
235	IFP Advisors, Inc	503	0.00%	$13K	+494 (+5488.9%)
236	Owl Creek Wealth Partners LLC	450	0.00%	$12K	—
237	HUNTINGTON NATIONAL BANK	445	0.00%	$11K	+25 (+6.0%)
238	ROTHSCHILD INVESTMENT LLC	437	0.00%	$11K	+149 (+51.7%)
239	Covestor Ltd	347	0.00%	$9K	+25 (+7.8%)
240	Steward Partners Investment Advisory, LLC	340	0.00%	$9K	+204 (+150.0%)
241	Private Wealth Management Group, LLC	296	0.00%	$8K	—
242	City State Bank	250	0.00%	$6K	—
243	Essential Partners LLC	236	0.00%	$6K	+191 (+424.4%)
244	SIGNATUREFD, LLC	230	0.00%	$6K	-11 (-4.6%)
245	SJS Investment Consulting Inc.	218	0.00%	$6K	+202 (+1262.5%)
246	EverSource Wealth Advisors, LLC	213	0.00%	$5K	-43 (-16.8%)
247	Parallel Advisors, LLC	213	0.00%	$5K	-145 (-40.5%)
248	HM PAYSON & CO	200	0.00%	$5K	NEW
249	Comprehensive Financial Planning, Inc./PA	200	0.00%	$5K	—
250	Parkside Financial Bank & Trust	189	0.00%	$5K	—

Exited since 2026-03-31 (31 filers; largest 25 shown)

Vestal Point Capital, LP	−1,773,326
STEMPOINT CAPITAL LP	−853,122
BRAIDWELL LP	−296,324
SCHRODER INVESTMENT MANAGEMENT GROUP	−285,820
Palo Alto Investors LP	−234,582
Balyasny Asset Management L.P.	−195,108
SECTOR GAMMA AS	−162,710
Y-Intercept (Hong Kong) Ltd	−78,007
POLEN CAPITAL MANAGEMENT LLC	−65,301
JANUS HENDERSON GROUP PLC	−60,028
Virtu Financial LLC	−43,362
Seven Fleet Capital Management LP	−35,286
JANE STREET GROUP, LLC	−32,147
TWINBEECH CAPITAL LP	−23,000
Amundi	−18,604
Perbak Capital Partners LLP	−13,175
Headlands Technologies LLC	−10,557
CWM, LLC	−4,651
Advisory Services Network, LLC	−848
QUADRANT CAPITAL GROUP LLC	−758
Quantbot Technologies LP	−512
NewEdge Advisors, LLC	−400
EASTERN BANK	−282
Aster Capital Management (DIFC) Ltd	−246
Root Financial Partners, LLC	−188

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting person	Form	Filed	Event	Shares	% of class
BAILLIE GIFFORD & CO	SCHEDULE 13G/A	2026-08-04	2026-06-30	11,960,230	7.5%
T. Rowe Price Investment Management, Inc.	SCHEDULE 13G/A	2026-07-08	2026-06-30	16,178,963	10.2%
Abigail P. Johnson	SCHEDULE 13G/A	2026-05-06	2026-03-31	6,672,999.32	4.2%
FMR LLC	SCHEDULE 13G/A	2026-05-06	2026-03-31	6,672,999.32	4.2%
The Vanguard Group	SCHEDULE 13G/A	2026-03-26	2026-03-13	0	0%
Wellington Group Holdings LLP	SCHEDULE 13G/A	2025-08-12	2025-06-30	5,664,733	3.9%
Wellington Investment Advisors Holdings LLP	SCHEDULE 13G/A	2025-08-12	2025-06-30	5,664,733	3.9%
Wellington Management Group LLP	SCHEDULE 13G/A	2025-08-12	2025-06-30	5,664,733	3.9%
BlackRock, Inc.	SCHEDULE 13G/A	2025-04-17	2025-03-31	10,490,610	7.2%
Capital Research Global Investors	SCHEDULE 13G/A	2025-02-13	2024-12-31	5,477,211	3.8%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.