Institutional HoldersBeta

Who holds a company, from every 13F filed for the quarter, one row per filer exactly as filed, against the latest share count the company reported.

Vaxcyte, Inc.

CIK 0001649094PCVX· quarter ending 2026-06-30

Shares outstanding

148,805,278

as of 2026-08-03 · 10-Q cover page, filed 2026-08-05

Held by 13F filers

167,366,414

112.5% of shares outstanding

Holders

332

-4 vs 2026-03-31 · 49 new, 53 exited

Value

$9.7B

shares +2.9% vs prior quarter

13F holdings exceed the share count. This is normal for heavily shorted stocks: shares lent to short sellers are reported by the lender and again by whoever bought them. Related filers occasionally report the same position twice as well.

Share classes (cover page of the 10-Q filed 2026-08-05, as of 2026-08-03)

Class	Listed	Shares outstanding	Held by 13F filers	% of class	Holders	CUSIP
Common stock	PCVX · NASDAQ	148,805,278	167,366,414	112.5%	332	92243G108

Each class is compared with its own share count; classes are not added together. A class that is not exchange-listed cannot be held by 13F filers.

Securities (CUSIPs held in 13F this quarter)

tick to combine

Count	CUSIP	Class (as filers write it)	Holders	Shares held	Class shares	% of class	Value	Options	Mapped via
	92243G108	COM	332	167,366,414	148,805,278	112.5%	$9.7B	11 call / 9 put	13DG	holders →

Totals combine the ticked CUSIPs and update as you tick: common-stock classes by default; preferreds, notes, warrants and depositary shares are listed but not counted. "holders →" shows a single CUSIP on its own. Bonds are held as a principal amount, not shares. Options positions are never counted as holdings.

Top holders (250 of 332 · 92243G108)

change vs 2026-03-31

#	Filer	Shares	% of class	Value	Change
1	FMR LLC	20,852,907	14.01%	$1.2B	+4,360,046 (+26.4%)
2	Jupiter Topco LLC	17,965,847	12.07%	$1.0B	NEW
3	T. Rowe Price Investment Management, Inc.	12,207,450	8.20%	$709.6M	+81,433 (+0.7%)
4	BlackRock, Inc.	12,068,129	8.11%	$701.5M	+1,096,313 (+10.0%)
5	RA CAPITAL MANAGEMENT, L.P.	11,474,599	7.71%	$667.0M	—
6	WELLINGTON MANAGEMENT GROUP LLP	7,001,565	4.71%	$407.0M	+1,386,968 (+24.7%)
7	VANGUARD PORTFOLIO MANAGEMENT LLC	6,593,551	4.43%	$383.3M	+314,006 (+5.0%)
8	VANGUARD CAPITAL MANAGEMENT LLC	6,348,679	4.27%	$369.0M	+81,978 (+1.3%)
9	STATE STREET CORP	5,351,698	3.60%	$311.1M	+911,179 (+20.5%)
10	Deep Track Capital, LP	4,300,000	2.89%	$250.0M	—
11	GEODE CAPITAL MANAGEMENT, LLC	3,459,363	2.32%	$201.1M	+258,080 (+8.1%)
12	PRICE T ROWE ASSOCIATES INC /MD/	2,988,999	2.01%	$173.8M	+2,012,570 (+206.1%)
13	Kynam Capital Management, LP	2,922,335	1.96%	$169.9M	—
14	WESTFIELD CAPITAL MANAGEMENT CO LP	2,472,428	1.66%	$143.7M	-1,173,710 (-32.2%)
15	Caligan Partners LP	2,188,636	1.47%	$127.2M	+127,917 (+6.2%)
16	Paradigm Biocapital Advisors LP	2,127,852	1.43%	$123.7M	-2,665,168 (-55.6%)
17	Bellevue Group AG	1,971,935	1.33%	$114.6M	+781,238 (+65.6%)
18	D. E. Shaw & Co., Inc.	1,901,872	1.28%	$110.6M	-963,446 (-33.6%)
19	MAVERICK CAPITAL LTD	1,613,242	1.08%	$93.8M	+235,504 (+17.1%)
20	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	1,464,812	0.98%	$85.1M	+60,864 (+4.3%)
21	Pivotal bioVenture Partners Investment Advisor LLC	1,425,470	0.96%	$82.9M	+53,045 (+3.9%)
22	EVENTIDE ASSET MANAGEMENT, LLC	1,380,219	0.93%	$80.2M	+5,927 (+0.4%)
23	GOLDMAN SACHS GROUP INC	1,350,760	0.91%	$78.5M	+75,724 (+5.9%)
24	NORTHERN TRUST CORP	1,280,172	0.86%	$74.4M	+135,703 (+11.9%)
25	NORGES BANK	1,245,805	0.84%	$72.4M	NEW
26	JPMORGAN CHASE & CO	1,227,346	0.82%	$71.2M	+219,032 (+21.7%)
27	DIMENSIONAL FUND ADVISORS LP	1,194,754	0.80%	$69.4M	+818,982 (+217.9%)
28	Polar Capital Holdings Plc	1,185,943	0.80%	$68.9M	-239,845 (-16.8%)
29	BANK OF AMERICA CORP /DE/	1,078,798	0.72%	$62.7M	-138,045 (-11.3%)
30	VANGUARD FIDUCIARY TRUST CO	930,642	0.63%	$54.1M	+88,369 (+10.5%)
31	ADAGE CAPITAL PARTNERS GP, L.L.C.	880,000	0.59%	$51.2M	+530,000 (+151.4%)
32	Saturn V Capital Management LP	852,914	0.57%	$49.6M	+28,382 (+3.4%)
33	Capital Research Global Investors	836,761	0.56%	$48.6M	-75,927 (-8.3%)
34	NOVO HOLDINGS A/S	800,000	0.54%	$46.5M	-60,000 (-7.0%)
35	MORGAN STANLEY	797,718	0.54%	$46.4M	+61,811 (+8.4%)
36	AMERICAN CENTURY COMPANIES INC	782,184	0.53%	$45.5M	+426,330 (+119.8%)
37	UBS Group AG	776,511	0.52%	$45.1M	-560,879 (-41.9%)
38	Clearbridge Investments, LLC	751,999	0.51%	$43.7M	+623,622 (+485.8%)
39	Pictet Asset Management Holding SA	694,719	0.47%	$40.4M	-5,592 (-0.8%)
40	Eversept Partners, LP	629,952	0.42%	$36.6M	-294,241 (-31.8%)
41	FEDERATED HERMES, INC.	601,161	0.40%	$34.9M	+34,333 (+6.1%)
42	BOOTHBAY FUND MANAGEMENT, LLC	534,163	0.36%	$31.1M	+12,144 (+2.3%)
43	ALGERT GLOBAL LLC	515,343	0.35%	$30.0M	+151,359 (+41.6%)
44	Bank of New York Mellon Corp	502,135	0.34%	$29.2M	+27,609 (+5.8%)
45	BROWN ADVISORY INC	487,017	0.33%	$28.3M	-24,684 (-4.8%)
46	JENNISON ASSOCIATES LLC	476,216	0.32%	$27.7M	-1,221,795 (-72.0%)
47	FRANKLIN RESOURCES INC	476,004	0.32%	$27.7M	+2,641 (+0.6%)
48	BNP PARIBAS FINANCIAL MARKETS	465,477	0.31%	$27.1M	+240,959 (+107.3%)
49	VICTORY CAPITAL MANAGEMENT INC	452,723	0.30%	$26.3M	-49,134 (-9.8%)
50	Elmind Capital, LP	452,340	0.30%	$26.3M	+135,840 (+42.9%)
51	Nantahala Capital Management, LLC	444,292	0.30%	$25.8M	+205,546 (+86.1%)
52	First Turn Management, LLC	443,836	0.30%	$25.8M	NEW
53	AMERIPRISE FINANCIAL INC	440,505	0.30%	$25.6M	+329,519 (+296.9%)
54	Ikarian Capital, LLC	388,682	0.26%	$22.6M	-40,000 (-9.3%)
55	Ensign Peak Advisors, Inc	381,686	0.26%	$22.2M	-64,217 (-14.4%)
56	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	371,550	0.25%	$21.6M	-39,055 (-9.5%)
57	RAYMOND JAMES FINANCIAL INC	369,406	0.25%	$21.5M	+109,011 (+41.9%)
58	Qube Research & Technologies Ltd	363,205	0.24%	$21.1M	+217,834 (+149.8%)
59	Aberdeen Group plc	362,756	0.24%	$21.1M	-616,213 (-62.9%)
60	Alyeska Investment Group, L.P.	362,272	0.24%	$21.1M	-73,309 (-16.8%)
61	First Light Asset Management, LLC	345,157	0.23%	$20.1M	+203,373 (+143.4%)
62	Boxer Capital Management, LLC	325,000	0.22%	$18.9M	—
63	Nuveen, LLC	324,762	0.22%	$18.9M	+3,973 (+1.2%)
64	Spruce Street Capital LP	306,047	0.21%	$17.8M	+60,338 (+24.6%)
65	Woodline Partners LP	274,740	0.18%	$16.0M	—
66	TD Asset Management Inc	274,640	0.18%	$16.0M	-38,122 (-12.2%)
67	TWO SIGMA INVESTMENTS, LP	273,876	0.18%	$15.9M	+162,566 (+146.0%)
68	Artisan Partners Limited Partnership	270,590	0.18%	$15.7M	-26,842 (-9.0%)
69	DEUTSCHE BANK AG\	258,338	0.17%	$15.0M	-41,355 (-13.8%)
70	Swiss National Bank	245,500	0.16%	$14.3M	+12,400 (+5.3%)
71	WCM INVESTMENT MANAGEMENT, LLC	243,502	0.16%	$14.1M	+130,871 (+116.2%)
72	University of Wisconsin Foundation	239,749	0.16%	$13.9M	—
73	California Public Employees Retirement System	212,472	0.14%	$12.4M	+60,917 (+40.2%)
74	Candriam S.C.A.	208,054	0.14%	$12.1M	-106,968 (-34.0%)
75	Assenagon Asset Management S.A.	202,866	0.14%	$11.8M	-961,947 (-82.6%)
76	CITIGROUP INC	202,673	0.14%	$11.8M	+70,495 (+53.3%)
77	Rosalind Advisors, Inc.	200,000	0.13%	$11.6M	-90,000 (-31.0%)
78	C WorldWide Group Holding A/S	195,000	0.13%	$11.3M	—
79	SEGALL BRYANT & HAMILL, LLC	187,269	0.13%	$10.9M	+3,334 (+1.8%)
80	Invesco Ltd.	173,439	0.12%	$10.1M	-10,604 (-5.8%)
81	RHUMBLINE ADVISERS	168,517	0.11%	$9.8M	+9,676 (+6.1%)
82	Russell Investments Group, Ltd.	164,334	0.11%	$9.6M	+126,155 (+330.4%)
83	ALLIANCEBERNSTEIN L.P.	159,909	0.11%	$9.3M	+22,827 (+16.7%)
84	READYSTATE ASSET MANAGEMENT LP	156,090	0.10%	$9.1M	+120,000 (+332.5%)
85	NEW YORK STATE COMMON RETIREMENT FUND	150,159	0.10%	$8.7M	+28,839 (+23.8%)
86	HighVista Strategies LLC	149,739	0.10%	$8.7M	+25,092 (+20.1%)
87	ADAR1 Capital Management, LLC	146,034	0.10%	$8.5M	-113,103 (-43.6%)
88	SILVERARC CAPITAL MANAGEMENT, LLC	145,000	0.10%	$8.4M	+19 (+0.0%)
89	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	140,833	0.09%	$8.2M	+4,733 (+3.5%)
90	MILLENNIUM MANAGEMENT LLC	136,625	0.09%	$7.9M	-157,070 (-53.5%)
91	Jump Financial, LLC	135,000	0.09%	$7.8M	NEW
92	WELLS FARGO & COMPANY/MN	132,513	0.09%	$7.7M	+79,407 (+149.5%)
93	Nicholas Investment Partners, LP	131,133	0.09%	$7.6M	NEW
94	Edgestream Partners, L.P.	131,115	0.09%	$7.6M	NEW
95	Trexquant Investment LP	128,333	0.09%	$7.5M	+79,121 (+160.8%)
96	NAN FUNG TRINITY (HK) LTD	122,930	0.08%	$7.1M	—
97	Creative Planning	120,771	0.08%	$7.0M	-19,406 (-13.8%)
98	FRED ALGER MANAGEMENT, LLC	116,965	0.08%	$6.8M	-153,150 (-56.7%)
99	VANGUARD ASSET MANAGEMENT, Ltd	115,020	0.08%	$6.7M	+5,777 (+5.3%)
100	MANUFACTURERS LIFE INSURANCE COMPANY, THE	108,504	0.07%	$6.3M	+22,138 (+25.6%)
101	NEXTBio Capital Management LP	107,223	0.07%	$6.2M	+12,000 (+12.6%)
102	Legal & General Group Plc	97,451	0.07%	$5.7M	+114 (+0.1%)
103	Ishara Investments LP	82,000	0.06%	$4.8M	+24,600 (+42.9%)
104	Man Group plc	81,632	0.05%	$4.7M	+62,375 (+323.9%)
105	MetLife Investment Management, LLC	76,628	0.05%	$4.5M	+8,774 (+12.9%)
106	STATE OF WISCONSIN INVESTMENT BOARD	71,772	0.05%	$4.2M	-1,130 (-1.6%)
107	ENVESTNET ASSET MANAGEMENT INC	70,581	0.05%	$4.1M	-27,574 (-28.1%)
108	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	67,741	0.05%	$3.9M	+3,915 (+6.1%)
109	SUPERSTRING CAPITAL MANAGEMENT LP	66,793	0.04%	$3.9M	-43,661 (-39.5%)
110	Universal- Beteiligungs- und Servicegesellschaft mbH	66,572	0.04%	$3.9M	+156 (+0.2%)
111	VOYA INVESTMENT MANAGEMENT LLC	65,110	0.04%	$3.8M	+3,547 (+5.8%)
112	Odyssean, LLC	62,638	0.04%	$3.6M	+42,527 (+211.5%)
113	Rafferty Asset Management, LLC	60,776	0.04%	$3.5M	-17,626 (-22.5%)
114	Rock Springs Capital Management LP	60,250	0.04%	$3.5M	—
115	SummitTX Capital, L.P.	57,355	0.04%	$3.3M	-17,487 (-23.4%)
116	Quantinno Capital Management LP	55,880	0.04%	$3.2M	+34,317 (+159.1%)
117	Zurcher Kantonalbank (Zurich Cantonalbank)	53,198	0.04%	$3.1M	+7,630 (+16.7%)
118	Bridgewater Associates, LP	50,217	0.03%	$2.9M	+37,314 (+289.2%)
119	Cormorant Asset Management, LP	50,000	0.03%	$2.9M	-50,000 (-50.0%)
120	Vanguard Global Advisers, LLC	49,787	0.03%	$2.9M	-1,863 (-3.6%)
121	Handelsbanken Fonder AB	49,400	0.03%	$2.9M	+8,500 (+20.8%)
122	DIADEMA PARTNERS LP	45,071	0.03%	$2.6M	+20,000 (+79.8%)
123	Hudson Bay Capital Management LP	45,000	0.03%	$2.6M	+14,030 (+45.3%)
124	BANQUE PICTET & CIE SA	44,736	0.03%	$2.6M	-17,110 (-27.7%)
125	ExodusPoint Capital Management, LP	42,357	0.03%	$2.5M	-83,470 (-66.3%)
126	TWINBEECH CAPITAL LP	42,192	0.03%	$2.5M	-23,708 (-36.0%)
127	LAZARD ASSET MANAGEMENT LLC	41,843	0.03%	$2.4M	-5,180 (-11.0%)
128	PRUDENTIAL FINANCIAL INC	41,570	0.03%	$2.4M	-17,755 (-29.9%)
129	State of New Jersey Common Pension Fund D	39,555	0.03%	$2.3M	-709 (-1.8%)
130	BARCLAYS PLC	39,053	0.03%	$2.3M	+7,065 (+22.1%)
131	GUGGENHEIM CAPITAL LLC	38,908	0.03%	$2.3M	+1,697 (+4.6%)
132	Police & Firemen's Retirement System of New Jersey	38,867	0.03%	$2.3M	+5,187 (+15.4%)
133	BROWN BROTHERS HARRIMAN & CO	38,033	0.03%	$2.2M	—
134	MARTINGALE ASSET MANAGEMENT L P	37,545	0.03%	$2.2M	+13,553 (+56.5%)
135	STRS OHIO	37,500	0.03%	$2.2M	+6,700 (+21.8%)
136	Corient Private Wealth LP	36,894	0.02%	$2.1M	+33,076 (+866.3%)
137	STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM	35,446	0.02%	$2.1M	+7,344 (+26.1%)
138	AQR CAPITAL MANAGEMENT LLC	34,563	0.02%	$2.0M	+9,105 (+35.8%)
139	Homestead Advisers Corp	34,400	0.02%	$2.0M	NEW
140	ARIZONA STATE RETIREMENT SYSTEM	33,755	0.02%	$2.0M	+1,394 (+4.3%)
141	SEI INVESTMENTS CO	33,516	0.02%	$1.9M	-29 (-0.1%)
142	Engineers Gate Manager LP	32,784	0.02%	$1.9M	NEW
143	PROFUND ADVISORS LLC	32,277	0.02%	$1.9M	-4,119 (-11.3%)
144	TEACHERS RETIREMENT SYSTEM OF THE STATE OF KENTUCKY	30,576	0.02%	$1.8M	+4,816 (+18.7%)
145	Vanguard Investments Australia, Ltd.	29,942	0.02%	$1.7M	—
146	Seven Fleet Capital Management LP	29,800	0.02%	$1.8M	-78,650 (-72.5%)
147	INTECH INVESTMENT MANAGEMENT LLC	28,577	0.02%	$1.7M	-2,636 (-8.4%)
148	KLP KAPITALFORVALTNING AS	27,700	0.02%	$1.6M	+1,800 (+6.9%)
149	SG Americas Securities, LLC	26,740	0.02%	$1.6M	-5,472 (-17.0%)
150	Vanguard Personalized Indexing Management, LLC	25,291	0.02%	$1.5M	-4,713 (-15.7%)
151	Mirae Asset Global Investments Co., Ltd.	24,214	0.02%	$1.4M	-1,049 (-4.2%)
152	Pathstone Holdings, LLC	23,876	0.02%	$1.4M	-3,290 (-12.1%)
153	China Universal Asset Management Co., Ltd.	22,981	0.02%	$1.3M	-2,559 (-10.0%)
154	ProShare Advisors LLC	21,990	0.01%	$1.3M	-2,459 (-10.1%)
155	Legato Capital Management LLC	21,645	0.01%	$1.3M	NEW
156	LPL Financial LLC	20,075	0.01%	$1.2M	-7,249 (-26.5%)
157	DekaBank Deutsche Girozentrale	20,000	0.01%	$1.1M	—
158	NOMURA ASSET MANAGEMENT INTERNATIONAL INC.	20,000	0.01%	$1.2M	—
159	RENAISSANCE TECHNOLOGIES LLC	19,634	0.01%	$1.1M	-35,100 (-64.1%)
160	XTX Topco Ltd	18,191	0.01%	$1.1M	+3,110 (+20.6%)
161	Financiere des Professionnels - Fonds d'investissement inc.	18,084	0.01%	$1.1M	-516 (-2.8%)
162	DNB Asset Management AS	18,041	0.01%	$1.0M	-1,059 (-5.5%)
163	New York Life Investment Management LLC	17,560	0.01%	$1.0M	NEW
164	Cinctive Capital Management LP	17,455	0.01%	$1.0M	-57,092 (-76.6%)
165	B. Metzler seel. Sohn & Co. AG	17,235	0.01%	$1.0M	—
166	GILDER GAGNON HOWE & CO LLC	17,064	0.01%	$992K	-111 (-0.6%)
167	Brevan Howard Capital Management LP	16,935	0.01%	$984K	+5,293 (+45.5%)
168	CIBC Bancorp USA Inc.	16,332	0.01%	$949K	+2,324 (+16.6%)
169	Persistent Asset Partners Ltd	15,105	0.01%	$878K	+9,786 (+184.0%)
170	CANADA LIFE ASSURANCE Co	14,463	0.01%	$838K	+1,060 (+7.9%)
171	ALLSTATE CORP	14,370	0.01%	$835K	-4,424 (-23.5%)
172	TEACHER RETIREMENT SYSTEM OF TEXAS	14,265	0.01%	$829K	-975 (-6.4%)
173	PUBLIC EMPLOYEES RETIREMENT ASSOCIATION OF COLORADO	14,024	0.01%	$815K	—
174	Aquatic Capital Management LLC	13,907	0.01%	$808K	NEW
175	Baird Financial Group, Inc.	13,813	0.01%	$803K	+25 (+0.2%)
176	Baker Avenue Asset Management, LP	13,279	0.01%	$772K	+136 (+1.0%)
177	XPONANCE LLC	13,050	0.01%	$759K	+1,652 (+14.5%)
178	J.Safra Asset Management Corp	12,564	0.01%	$730K	+1,714 (+15.8%)
179	LANDSCAPE CAPITAL MANAGEMENT, L.L.C.	12,211	0.01%	$710K	NEW
180	Marex Group Ltd	12,127	0.01%	$705K	+2,521 (+26.2%)
181	Ameritas Investment Partners, Inc.	11,960	0.01%	$695K	+1,676 (+16.3%)
182	PNC Financial Services Group, Inc.	11,800	0.01%	$686K	-3,156 (-21.1%)
183	Thrivent Financial for Lutherans	11,742	0.01%	$683K	—
184	SUSQUEHANNA INTERNATIONAL GROUP, LLP	11,206	0.01%	$651K	-35,867 (-76.2%)
185	Graham Capital Management, L.P.	11,107	0.01%	$646K	NEW
186	ROYAL BANK OF CANADA	10,734	0.01%	$625K	+2,183 (+25.5%)
187	GROUP ONE TRADING LLC	10,632	0.01%	$618K	NEW
188	Federation des caisses Desjardins du Quebec	10,382	0.01%	$604K	+10,190 (+5307.3%)
189	Cetera Investment Advisers	10,362	0.01%	$602K	+767 (+8.0%)
190	Rockefeller Capital Management L.P.	10,133	0.01%	$589K	+254 (+2.6%)
191	Quarry LP	10,043	0.01%	$584K	+3,043 (+43.5%)
192	Amundi	10,003	0.01%	$581K	-2,955 (-22.8%)
193	Mariner, LLC	9,545	0.01%	$555K	-3,435 (-26.5%)
194	Daiwa Securities Group Inc.	9,114	0.01%	$530K	-230 (-2.5%)
195	E Fund Management Co., Ltd.	8,898	0.01%	$517K	-1,201 (-11.9%)
196	Vident Advisory, LLC	8,361	0.01%	$486K	-1,143 (-12.0%)
197	OSAIC HOLDINGS, INC.	8,028	0.01%	$467K	-842 (-9.5%)
198	CANADA PENSION PLAN INVESTMENT BOARD	7,900	0.01%	$459K	-3,800 (-32.5%)
199	Allianz Asset Management GmbH	7,891	0.01%	$459K	+791 (+11.1%)
200	Swiss Life Asset Management Ltd	7,818	0.01%	$454K	+935 (+13.6%)
201	Meeder Advisory Services, Inc.	7,785	0.01%	$453K	-550 (-6.6%)
202	PRINCIPAL FINANCIAL GROUP INC	7,711	0.01%	$448K	+2,436 (+46.2%)
203	Allspring Global Investments Holdings, LLC	7,364	0.00%	$413K	+895 (+13.8%)
204	Sumitomo Mitsui Trust Group, Inc.	7,304	0.00%	$425K	-27 (-0.4%)
205	Marks Group Wealth Management, Inc	7,224	0.00%	$420K	+666 (+10.2%)
206	Virtus Investment Advisers, LLC	6,894	0.00%	$401K	+768 (+12.5%)
207	Values First Advisors, Inc.	6,768	0.00%	$393K	+151 (+2.3%)
208	SIGNATUREFD, LLC	6,510	0.00%	$378K	-610 (-8.6%)
209	JANE STREET GROUP, LLC	6,401	0.00%	$372K	-246,476 (-97.5%)
210	JONES FINANCIAL COMPANIES LLLP	6,068	0.00%	$343K	+2,497 (+69.9%)
211	Avior Wealth Management, LLC	5,744	0.00%	$334K	-2,903 (-33.6%)
212	Sanctuary Advisors, LLC	5,596	0.00%	$325K	-668 (-10.7%)
213	MML INVESTORS SERVICES, LLC	5,559	0.00%	$323K	NEW
214	Empowered Funds, LLC	5,424	0.00%	$315K	-4,268 (-44.0%)
215	Tower Research Capital LLC (TRC)	4,988	0.00%	$290K	-7,172 (-59.0%)
216	Nykredit A/S	4,823	0.00%	$280K	NEW
217	Caprock Group, LLC	4,798	0.00%	$279K	NEW
218	Cerity Partners LLC	4,722	0.00%	$274K	-908 (-16.1%)
219	CAPSTONE INVESTMENT ADVISORS, LLC	4,666	0.00%	$271K	NEW
220	NEUBERGER BERMAN GROUP LLC	4,593	0.00%	$267K	-319 (-6.5%)
221	CIBC Asset Management Inc	4,516	0.00%	$263K	—
222	O'SHAUGHNESSY ASSET MANAGEMENT, LLC	4,371	0.00%	$254K	NEW
223	M&T Bank Corp	4,275	0.00%	$248K	+35 (+0.8%)
224	FOX RUN MANAGEMENT, L.L.C.	4,248	0.00%	$247K	NEW
225	Callan Family Office, LLC	4,199	0.00%	$244K	+10 (+0.2%)
226	HSBC HOLDINGS PLC	4,151	0.00%	$241K	+159 (+4.0%)
227	NISA INVESTMENT ADVISORS, LLC	3,980	0.00%	$231K	+110 (+2.8%)
228	Blueshift Asset Management, LLC	3,934	0.00%	$229K	NEW
229	KBC Group NV	3,898	0.00%	$227K	—
230	Bayforest Capital Ltd	3,706	0.00%	$215K	+3,359 (+968.0%)
231	Storgate, LLC	3,591	0.00%	$209K	NEW
232	AMALGAMATED BANK	3,576	0.00%	$208K	-21 (-0.6%)
233	Mitsubishi UFJ Trust & Banking Corp	3,146	0.00%	$183K	NEW
234	GF FUND MANAGEMENT CO. LTD.	3,048	0.00%	$177K	+108 (+3.7%)
235	Parallel Advisors, LLC	2,260	0.00%	$131K	+566 (+33.4%)
236	FNY Investment Advisers, LLC	2,000	0.00%	$116K	—
237	BNP PARIBAS ASSET MANAGEMENT Holding S.A.	1,961	0.00%	$114K	-248 (-11.2%)
238	Walleye Capital LLC	1,942	0.00%	$113K	-48,708 (-96.2%)
239	Advisors Asset Management, Inc.	1,921	0.00%	$112K	+24 (+1.3%)
240	ASSETMARK, INC	1,851	0.00%	$108K	-28 (-1.5%)
241	US BANCORP \DE\	1,502	0.00%	$87K	-140 (-8.5%)
242	Eurizon Capital SGR S.p.A.	1,383	0.00%	$80K	—
243	FIFTH THIRD BANCORP	1,379	0.00%	$80K	-826 (-37.5%)
244	GAMMA Investing LLC	1,347	0.00%	$78K	+66 (+5.2%)
245	NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO	1,344	0.00%	$78K	+291 (+27.6%)
246	Kemnay Advisory Services Inc.	1,299	0.00%	$76K	+119 (+10.1%)
247	Allworth Financial LP	1,285	0.00%	$75K	-152 (-10.6%)
248	Sterling Capital Management LLC	1,174	0.00%	$68K	-2,965 (-71.6%)
249	Pacer Advisors, Inc.	1,015	0.00%	$59K	NEW
250	Capital Advisors, Ltd. LLC	1,000	0.00%	$58K	—

Exited since 2026-03-31 (53 filers; largest 25 shown)

JANUS HENDERSON GROUP PLC	−16,575,875
FARALLON CAPITAL MANAGEMENT LLC	−1,261,000
BAUPOST GROUP LLC/MA	−800,000
Trails Edge Capital Partners, LP	−662,470
Jones Hill Capital LP	−644,831
Point72 Asset Management, L.P.	−620,401
Balyasny Asset Management L.P.	−514,577
Soleus Capital Management, L.P.	−385,472
Logos Global Management LP	−300,000
Freestone Grove Partners LP	−224,731
STEMPOINT CAPITAL LP	−118,122
Sofinnova Investments, Inc.	−50,600
Burkehill Global Management, LP	−50,000
ArrowMark Colorado Holdings LLC	−40,000
HRT FINANCIAL LP	−37,318
Ghisallo Capital Management LLC	−25,000
Progeny 3, Inc.	−23,300
CIBC WORLD MARKET INC.	−20,516
DV Trading LLC	−20,000
Brooklyn Investment Group	−9,370
VARCOV Co.	−8,329
Abel Hall, LLC	−6,463
Dynamic Technology Lab Private Ltd	−6,110
INCEPTIONR LLC	−5,753
Atom Investors LP	−5,653

Large holders from 13D/13G (latest per reporting person, last 2 years)

Filed when a holder crosses 5% or changes materially, so often more current than the quarter-end 13F. Percentages are the filer's own, against the share count they used.

Reporting person	Form	Filed	Event	Shares	% of class
JANUS HENDERSON GROUP Ltd.	SCHEDULE 13G/A	2026-08-11	2026-07-31	18,040,375	12.1%
Abigail P. Johnson	SCHEDULE 13G/A	2026-08-06	2026-06-30	20,798,906.96	14.4%
FMR LLC	SCHEDULE 13G/A	2026-08-06	2026-06-30	20,798,906.96	14.4%
JANUS HENDERSON GROUP PLC	SCHEDULE 13G/A	2026-05-15	2026-03-31	16,575,875	11.5%
Peter Kolchinsky	SCHEDULE 13G/A	2026-05-15	2026-03-31	12,974,599	8.9%
RA Capital Healthcare Fund, L.P.	SCHEDULE 13G/A	2026-05-15	2026-03-31	12,699,089	8.7%
RA Capital Management, L.P.	SCHEDULE 13G/A	2026-05-15	2026-03-31	12,974,599	8.9%
Rajeev Shah	SCHEDULE 13G/A	2026-05-15	2026-03-31	12,974,599	8.9%
Vanguard Capital Management	SCHEDULE 13G	2026-04-30	2026-03-31	7,252,053	5.03%
The Vanguard Group	SCHEDULE 13G/A	2026-03-27	2026-03-13	0	0%
T. Rowe Price Investment Management, Inc.	SCHEDULE 13G/A	2026-02-17	2025-12-31	11,296,333	8.6%
BlackRock, Inc.	SCHEDULE 13G/A	2025-04-17	2025-03-31	9,825,638	7.6%

Source: SEC Form 13F-HR information tables for the quarter ending 2026-06-30 (filed within 45 days of quarter end; amendments applied; values normalised to dollars), the share count from the company's latest 10-K/10-Q XBRL cover page, and Schedule 13D/13G filings. Each row is a legal 13F filer as filed — related entities (e.g. Vanguard's advisers) are not merged. Retail brokerage accounts are not reported on 13F, so the remainder is mostly retail and insiders.